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Agriculture is one of the most important human activities providing food and more agricultural goods for seven billion people around the world and is of special importance in sub-Saharan Africa. The majority of people depends on the agricultural sector for their livelihoods and will suffer from negative climate change impacts on agriculture until the middle and end of the 21st century, even more if weak governments, economic crises or violent conflicts endanger the countries’ food security. The impact of temperature increases and changing precipitation patterns on agricultural vegetation motivated this thesis in the first place. Analyzing the potentials of reducing negative climate change impacts by adapting crop management to changing climate is a second objective of the thesis. As a precondition for simulating climate change impacts on agricultural crops with a global crop model first the timing of sowing in the tropics was improved and validated as this is an important factor determining the length and timing of the crops´ development phases, the occurrence of water stress and final crop yield. Crop yields are projected to decline in most regions which is evident from the results of this thesis, but the uncertainties that exist in climate projections and in the efficiency of adaptation options because of political, economical or institutional obstacles have to be considered. The effect of temperature increases and changing precipitation patterns on crop yields can be analyzed separately and varies in space across the continent. Southern Africa is clearly the region most susceptible to climate change, especially to precipitation changes. The Sahel north of 13° N and parts of Eastern Africa with short growing seasons below 120 days and limited wet season precipitation of less than 500 mm are also vulnerable to precipitation changes while in most other part of East and Central Africa, in contrast, the effect of temperature increase on crops overbalances the precipitation effect and is most pronounced in a band stretching from Angola to Ethiopia in the 2060s. The results of this thesis confirm the findings from previous studies on the magnitude of climate change impact on crops in sub-Saharan Africa but beyond that helps to understand the drivers of these changes and the potential of certain management strategies for adaptation in more detail. Crop yield changes depend on the initial growing conditions, on the magnitude of climate change, and on the crop, cropping system and adaptive capacity of African farmers which is only now evident from this comprehensive study for sub-Saharan Africa. Furthermore this study improves the representation of tropical cropping systems in a global crop model and considers the major food crops cultivated in sub-Saharan Africa and climate change impacts throughout the continent.
Rainfall, snow-, and glacial melt throughout the Himalaya control river discharge, which is vital for maintaining agriculture, drinking water and hydropower generation. However, the spatiotemporal contribution of these discharge components to Himalayan rivers is not well understood, mainly because of the scarcity of ground-based observations. Consequently, there is also little known about the triggers and sources of peak sediment flux events, which account for extensive hydropower reservoir filling and turbine abrasion. We therefore lack basic information on the distribution of water resources and controls of erosion processes. In this thesis, I employ various methods to assess and quantify general characteristics of and links between precipitation, river discharge, and sediment flux in the Sutlej Valley. First, I analyze daily precipitation data (1998-2007) from 80 weather stations in the western Himalaya, to decipher the distribution of rain- and snowfall. Rainfall magnitude frequency analyses indicate that 40% of the summer rainfall budget is attributed to monsoonal rainstorms, which show higher variability in the orogenic interior than in frontal regions. Combined analysis of rainstorms and sediment flux data of a major Sutlej River tributary indicate that monsoonal rainfall has a first order control on erosion processes in the orogenic interior, despite the dominance of snowfall in this region. Second, I examine the contribution of rainfall, snow and glacial melt to river discharge in the Sutlej Valley (s55,000 km2), based on a distributed hydrological model, which covers the period 2000-2008. To achieve high spatial and daily resolution despite limited ground-based observations the hydrological model is forced by daily remote sensing data, which I adjusted and calibrated with ground station data. The calibration shows that the Tropical Rainfall Measuring Mission (TRMM) 3B42 rainfall product systematically overestimates rainfall in semi-arid and arid regions, increasing with aridity. The model results indicate that snowmelt-derived discharge (74%) is most important during the pre-monsoon season (April to June) whereas rainfall (56%) and glacial melt (17%) dominate the monsoon season (July-September). Therefore, climate change most likely causes a reduction in river discharge during the pre-monsoon season, which especially affects the orogenic interior. Third, I investigate the controls on suspended sediment flux in different parts of the Sutlej catchments, based on daily gauging data from the past decade. In conjunction with meteorological data, earthquake records, and rock strength measurements I find that rainstorms are the most frequent trigger of high-discharge events with peaks in suspended sediment concentrations (SSC) that account for the bulk of the suspended sediment flux. The suspended sediment flux increases downstream, mainly due to increases in runoff. Pronounced erosion along the Himalayan Front occurs throughout the monsoon season, whereas efficient erosion of the orogenic interior is confined to single extreme events. The results of this thesis highlight the importance of snow and glacially derived melt waters in the western Himalaya, where extensive regions receive only limited amounts of monsoonal rainfall. These regions are therefore particularly susceptible to global warming with major implications on the hydrological cycle. However, the sediment discharge data show that infrequent monsoonal rainstorms that pass the orographic barrier of the Higher Himalaya are still the primary trigger of the highest-impact erosion events, despite being subordinate to snow and glacially–derived discharge. These findings may help to predict peak sediment flux events and could underpin the strategic development of preventative measures for hydropower infrastructures.
Species respond to environmental change by dynamically adjusting their geographical ranges. Robust predictions of these changes are prerequisites to inform dynamic and sustainable conservation strategies. Correlative species distribution models (SDMs) relate species’ occurrence records to prevailing environmental factors to describe the environmental niche. They have been widely applied in global change context as they have comparably low data requirements and allow for rapid assessments of potential future species’ distributions. However, due to their static nature, transient responses to environmental change are essentially ignored in SDMs. Furthermore, neither dispersal nor demographic processes and biotic interactions are explicitly incorporated. Therefore, it has often been suggested to link statistical and mechanistic modelling approaches in order to make more realistic predictions of species’ distributions for scenarios of environmental change. In this thesis, I present two different ways of such linkage. (i) Mechanistic modelling can act as virtual playground for testing statistical models and allows extensive exploration of specific questions. I promote this ‘virtual ecologist’ approach as a powerful evaluation framework for testing sampling protocols, analyses and modelling tools. Also, I employ such an approach to systematically assess the effects of transient dynamics and ecological properties and processes on the prediction accuracy of SDMs for climate change projections. That way, relevant mechanisms are identified that shape the species’ response to altered environmental conditions and which should hence be considered when trying to project species’ distribution through time. (ii) I supplement SDM projections of potential future habitat for black grouse in Switzerland with an individual-based population model. By explicitly considering complex interactions between habitat availability and demographic processes, this allows for a more direct assessment of expected population response to environmental change and associated extinction risks. However, predictions were highly variable across simulations emphasising the need for principal evaluation tools like sensitivity analysis to assess uncertainty and robustness in dynamic range predictions. Furthermore, I identify data coverage of the environmental niche as a likely cause for contrasted range predictions between SDM algorithms. SDMs may fail to make reliable predictions for truncated and edge niches, meaning that portions of the niche are not represented in the data or niche edges coincide with data limits. Overall, my thesis contributes to an improved understanding of uncertainty factors in predictions of range dynamics and presents ways how to deal with these. Finally I provide preliminary guidelines for predictive modelling of dynamic species’ response to environmental change, identify key challenges for future research and discuss emerging developments.
In den letzten drei Jahrzehnten wurden in einigen Seen und Feuchtgebieten in bewaldeten Einzugsgebieten Nordost-Brandenburgs sinkende Wasserstände beobachtet. In diesen Gebieten bestimmt die Grundwasserneubildung im Einzugsgebiet maßgeblich das Wasserdargebot der Seen und Feuchtgebiete, die deshalb hier als grundwasserabhängige Landschaftselemente bezeichnet werden. Somit weisen die sinkenden Wasserstände auf einen Rückgang der wegen des geringen Niederschlagsdargebotes ohnehin schon geringen Grundwasserneubildung hin. Die Höhe der Grundwasserneubildung ist neben den hydroklimatischen Randbedingungen auch von der Landnutzung abhängig. Veränderungen in der Waldvegetation und der hydroklimatischen Randbedingungen bewirken Änderungen der Grundwasserneubildung und beeinflussen somit auch den Wasserhaushalt der Seen und Feuchtgebiete. Aktuell wird die Waldvegetation durch Kiefernmonokulturen dominiert, mit im Vergleich zu anderen Baumarten höherer Evapotranspiration. Entwicklungen in der Forstwirtschaft streben die Verringerung von Kiefernmonokulturen an. Diese sollen langfristig auf geeigneten Standorten durch Laubmischwälder ersetzt werden. Dadurch lassen sich eine geringere Evapotranspiration und damit eine höhere Grundwasserneubildung erreichen. In der vorliegenden Arbeit werden am Beispiel des Redernswalder Sees und des Briesensees die Ursachen der beobachteten sinkenden Wasserstände analysiert. Ihre Wasserstände nahmen in den letzten 25 Jahren um mehr als 3 Meter ab. Weiterhin wird untersucht, wie die erwarteten Klimaänderungen und Veränderungen in der Waldbewirtschaftung die zukünftige Grundwasserneubildung und den Wasserhaushalt von Seen beeinflussen können. Die Entwicklung der Grundwasserneubildung im Untersuchungsgebiet wurde mit dem Wasserhaushaltsmodell WaSiM-ETH simuliert. Die Analyse der Wechselwirkungen der Seen mit dem regionalen quartären Grundwasserleitersystem erfolgte mit dem 3D-Grundwassermodell FEFLOW. Mögliche zukünftige Veränderungen der Grundwasserneubildung und der Seewasserstände durch Klimaänderungen und Waldumbau wurden mit Szenarienrechnungen bis zum Jahr 2100 analysiert. Die modellgestützte Analyse zeigte, dass die beobachteten abnehmenden Wasserstände zu etwa gleichen Anteilen durch Veränderungen der hydroklimatischen Randbedingungen sowie durch Veränderungen in der Waldvegetation und damit abnehmenden Grundwasserneubildungsraten zu erklären sind. Die zukünftigen Entwicklungen der Grundwasserneubildung und der Wasserstände sind geprägt von sich ändernden hydroklimatischen Randbedingungen und einem sukzessiven Wandel der Kiefernbestände zu Laubwäldern. Der Waldumbau hat positive Wirkungen auf die Grundwasserneubildung und damit auf die Wasserstände. Damit können die Einflüsse des eingesetzten REMO-A1B-Klimaszenarios zum Ende des Modellzeitraumes durch den Waldumbau nicht kompensiert werden, das Sinken des Wasserstandes wird jedoch wesentlich reduziert. Bei dem moderateren REMO-B1-Klimaszenario werden die Wasserstände des Jahres 2008 durch den Waldumbau bis zum Jahr 2100 überschritten.
The Greenland Ice Sheet (GIS) contains enough water volume to raise global sea level by over 7 meters. It is a relic of past glacial climates that could be strongly affected by a warming world. Several studies have been performed to investigate the sensitivity of the ice sheet to changes in climate, but large uncertainties in its long-term response still exist. In this thesis, a new approach has been developed and applied to modeling the GIS response to climate change. The advantages compared to previous approaches are (i) that it can be applied over a wide range of climatic scenarios (both in the deep past and the future), (ii) that it includes the relevant feedback processes between the climate and the ice sheet and (iii) that it is highly computationally efficient, allowing simulations over very long timescales. The new regional energy-moisture balance model (REMBO) has been developed to model the climate and surface mass balance over Greenland and it represents an improvement compared to conventional approaches in modeling present-day conditions. Furthermore, the evolution of the GIS has been simulated over the last glacial cycle using an ensemble of model versions. The model performance has been validated against field observations of the present-day climate and surface mass balance, as well as paleo information from ice cores. The GIS contribution to sea level rise during the last interglacial is estimated to be between 0.5-4.1 m, consistent with previous estimates. The ensemble of model versions has been constrained to those that are consistent with the data, and a range of valid parameter values has been defined, allowing quantification of the uncertainty and sensitivity of the modeling approach. Using the constrained model ensemble, the sensitivity of the GIS to long-term climate change was investigated. It was found that the GIS exhibits hysteresis behavior (i.e., it is multi-stable under certain conditions), and that a temperature threshold exists above which the ice sheet transitions to an essentially ice-free state. The threshold in the global temperature is estimated to be in the range of 1.3-2.3°C above preindustrial conditions, significantly lower than previously believed. The timescale of total melt scales non-linearly with the overshoot above the temperature threshold, such that a 2°C anomaly causes the ice sheet to melt in ca. 50,000 years, but an anomaly of 6°C will melt the ice sheet in less than 4,000 years. The meltback of the ice sheet was found to become irreversible after a fraction of the ice sheet is already lost – but this level of irreversibility also depends on the temperature anomaly.
Lake ecosystems across the globe have responded to climate warming of recent decades. However, correctly attributing observed changes to altered climatic conditions is complicated by multiple anthropogenic influences on lakes. This thesis contributes to a better understanding of climate impacts on freshwater phytoplankton, which forms the basis of the food chain and decisively influences water quality. The analyses were, for the most part, based on a long-term data set of physical, chemical and biological variables of a shallow, polymictic lake in north-eastern Germany (Müggelsee), which was subject to a simultaneous change in climate and trophic state during the past three decades. Data analysis included constructing a dynamic simulation model, implementing a genetic algorithm to parameterize models, and applying statistical techniques of classification tree and time-series analysis. Model results indicated that climatic factors and trophic state interactively determine the timing of the phytoplankton spring bloom (phenology) in shallow lakes. Under equally mild spring conditions, the phytoplankton spring bloom collapsed earlier under high than under low nutrient availability, due to a switch from a bottom-up driven to a top-down driven collapse. A novel approach to model phenology proved useful to assess the timings of population peaks in an artificially forced zooplankton-phytoplankton system. Mimicking climate warming by lengthening the growing period advanced algal blooms and consequently also peaks in zooplankton abundance. Investigating the reasons for the contrasting development of cyanobacteria during two recent summer heat wave events revealed that anomalously hot weather did not always, as often hypothesized, promote cyanobacteria in the nutrient-rich lake studied. The seasonal timing and duration of heat waves determined whether critical thresholds of thermal stratification, decisive for cyanobacterial bloom formation, were crossed. In addition, the temporal patterns of heat wave events influenced the summer abundance of some zooplankton species, which as predators may serve as a buffer by suppressing phytoplankton bloom formation. This thesis adds to the growing body of evidence that lake ecosystems have strongly responded to climatic changes of recent decades. It reaches beyond many previous studies of climate impacts on lakes by focusing on underlying mechanisms and explicitly considering multiple environmental changes. Key findings show that climate impacts are more severe in nutrient-rich than in nutrient-poor lakes. Hence, to develop lake management plans for the future, limnologists need to seek a comprehensive, mechanistic understanding of overlapping effects of the multi-faceted human footprint on aquatic ecosystems.
Forests are a key resource serving a multitude of functions such as providing income to forest owners, supplying industries with timber, protecting water resources, and maintaining biodiversity. Recently much attention has been given to the role of forests in the global carbon cycle and their management for increased carbon sequestration as a possible mitigation option against climate change. Furthermore, the use of harvested wood can contribute to the reduction of atmospheric carbon through (i) carbon sequestration in wood products, (ii) the substitution of non-wood products with wood products, and (iii) through the use of wood as a biofuel to replace fossil fuels. Forest resource managers are challenged by the task to balance these multiple while simultaneously meeting economic requirements and taking into consideration the demands of stakeholder groups. Additionally, risks and uncertainties with regard to uncontrollable external variables such as climate have to be considered in the decision making process. In this study a scientific stakeholder dialogue with forest-related stakeholder groups in the Federal State of Brandenburg was accomplished. The main results of this dialogue were the definition of major forest functions (carbon sequestration, groundwater recharge, biodiversity, and timber production) and priority setting among them by the stakeholders using the pair-wise comparison technique. The impact of different forest management strategies and climate change scenarios on the main functions of forest ecosystems were evaluated at the Kleinsee management unit in south-east Brandenburg. Forest management strategies were simulated over 100 years using the forest growth model 4C and a wood product model (WPM). A current climate scenario and two climate change scenarios based on global circulation models (GCMs) HadCM2 and ECHAM4 were applied. The climate change scenario positively influenced stand productivity, carbon sequestration, and income. The impact on the other forest functions was small. Furthermore, the overall utility of forest management strategies were compared under the priority settings of stakeholders by a multi-criteria analysis (MCA) method. Significant differences in priority setting and the choice of an adequate management strategy were found for the environmentalists on one side and the more economy-oriented forest managers of public and private owned forests on the other side. From an ecological perspective, a conservation strategy would be preferable under all climate scenarios, but the business as usual management would also fit the expectations under the current climate. In contrast, a forest manager in public-owned forests or a private forest owner would prefer a management strategy with an intermediate thinning intensity and a high share of pine stands to enhance income from timber production while maintaining the other forest functions. The analysis served as an example for the combined application of simulation tools and a MCA method for the evaluation of management strategies under multi-purpose and multi-user settings with changing climatic conditions. Another focus was set on quantifying the overall effect of forest management on carbon sequestration in the forest sector and the wood industry sector plus substitution effects. To achieve this objective, the carbon emission reduction potential of material and energy substitution (Smat and Sen) was estimated based on a literature review. On average, for each tonne of dry wood used in a wood product substituting a non-wood product, 0.71 fewer tonnes of fossil carbon are emitted into to the atmosphere. Based on Smat and Sen, the calculation of the carbon emission reduction through substitution was implemented in the WPM. Carbon sequestration and substitution effects of management strategies were simulated at three local scales using the WPM and the forest growth models 4C (management unit level) or EFISCEN (federal state of Brandenburg and Germany). An investigation was conducted on the influence of uncertainties in the initialisation of the WPM, Smat, and basic conditions of the wood product sector on carbon sequestration. Results showed that carbon sequestration in the wood industry sector plus substitution effects exceeded sequestration in the forest sector. In contrast to the carbon pools in the forest sector, which acted as sink or source, the substitution effects continually reduced carbon emission as long as forests are managed and timber is harvested. The main climate protection function was investigated for energy substitution which accounted for about half of the total carbon sequestration, followed by carbon storage in landfills. In Germany, the absolute annual carbon sequestration in the forest and wood industry sector plus substitution effects was 19.9 Mt C. Over 50 years the wood industry sector contributed 70% of the total carbon sequestration plus substitution effects.
The need to develop sustainable resource management strategies for semi-arid and arid rangelands is acute as non-adapted grazing strategies lead to irreversible environmental problems such as desertification and associated loss of economic support to society. In such vulnerable ecosystems, successful implementation of sustainable management strategies depends on well-founded under-standing of processes at different scales that underlay the complex system dynamic. There is ample evidence that, in contrast to traditional sectoral approaches, only interdisciplinary research does work for resolving problems in conservation and natural resource management. In this thesis I combined a range of modeling approaches that integrate different disciplines and spatial scales in order to contribute to basic guidelines for sustainable management of semi-arid and arid range-lands. Since water availability and livestock management are seen as most potent determinants for the dynamics of semi-arid and arid ecosystems I focused on (i) the interaction of ecological and hydro-logical processes and (ii) the effect of farming strategies. First, I developed a grid-based and small-scaled model simulating vegetation dynamics and inter-linked hydrological processes. The simulation results suggest that ecohydrological interactions gain importance in rangelands with ascending slope where vegetation cover serves to obstruct run-off and decreases evaporation from the soil. Disturbances like overgrazing influence these positive feedback mechanisms by affecting vegetation cover and composition. In the second part, I present a modeling approach that has the power to transfer and integrate ecological information from the small scale vegetation model to the landscape scale, most relevant for the conservation of biodiversity and sustainable management of natural resources. I combined techniques of stochastic modeling with remotely sensed data and GIS to investigate to which ex-tent spatial interactions, like the movement of surface water by run-off in water limited environments, affect ecosystem functioning at the landscape scale. My simulation experiments show that overgrazing decreases the number of vegetation patches that act as hydrological sinks and run-off increases. The results of both simulation models implicate that different vegetation types should not only be regarded as provider of forage production but also as regulator of ecosystem functioning. Vegetation patches with good cover of perennial vegetation are capable to catch and conserve surface run-off from degraded surrounding areas. Therefore, downstream out of the simulated system is prevented and efficient use of water resources is guaranteed at all times. This consequence also applies to commercial rotational grazing strategies for semi-arid and arid rangelands with ascending slope where non-degraded paddocks act as hydrological sinks. Finally, by the help of an integrated ecological-economic modeling approach, I analyzed the relevance of farmers’ ecological knowledge for longterm functioning of semi-arid and arid grazing systems under current and future climatic conditions. The modeling approach consists of an ecological and an economic module and combines relevant processes on either level. Again, vegetation dynamics and forage productivity is derived by the small-scaled vegetation model. I showed that sustainable management of semi-arid and arid rangelands relies strongly on the farmers’ knowledge on how the ecosystem works. Furthermore, my simulation results indicate that the projected lower annual rainfall due to climate change in combination with non-adapted grazing strategies adds an additional layer of risk to these ecosystems that are already prone to land degradation. All simulation models focus on the most essential factors and ignore specific details. Therefore, even though all simulation models are parameterized for a specific dwarf shrub savanna in arid southern Namibia, the conclusions drawn are applicable for semi-arid and arid rangelands in general.
Soils contain a large amount of carbon (C) that is a critical regulator of the global C budget. Already small changes in the processes governing soil C cycling have the potential to release considerable amounts of CO2, a greenhouse gas (GHG), adding additional radiative forcing to the atmosphere and hence to changing climate. Increased temperatures will probably create a feedback, causing soils to release more GHGs. Furthermore changes in soil C balance impact soil fertility and soil quality, potentially degrading soils and reducing soils function as important resource. Consequently the assessment of soil C dynamics under present, recent past and future environmental conditions is not only of scientific interest and requires an integrated consideration of main factors and processes governing soil C dynamics. To perform this assessment an eco-hydrological modelling tool was used and extended by a process-based description of coupled soil carbon and nitrogen turnover. The extended model aims at delivering sound information on soil C storage changes beside changes in water quality, quantity and vegetation growth under global change impacts in meso- to macro-scale river basins, exemplary demonstrated for a Central European river basin (the Elbe). As a result this study: ▪ Provides information on joint effects of land-use (land cover and land management) and climate changes on croplands soil C balance in the Elbe river basin (Central Europe) presently and in the future. ▪ Evaluates which processes, and at what level of process detail, have to be considered to perform an integrated simulation of soil C dynamics at the meso- to macro-scale and demonstrates the model’s capability to simulate these processes compared to observations. ▪ Proposes a process description relating soil C pools and turnover properties to readily measurable quantities. This reduces the number of model parameters, enhances the comparability of model results to observations, and delivers same performance simulating long-term soil C dynamics as other models. ▪ Presents an extensive assessment of the parameter and input data uncertainty and their importance both temporally and spatially on modelling soil C dynamics. For the basin scale assessments it is estimated that croplands in the Elbe basin currently act as a net source of carbon (net annual C flux of 11 g C m-2 yr-1, 1.57 106 tons CO2 yr-1 entire croplands on average). Although this highly depends on the amount of harvest by-products remaining on the field. Future anticipated climate change and observed climate change in the basin already accelerates soil C loss and increases source strengths (additional 3.2 g C m-2 yr-1, 0.48 106 tons CO2 yr-1 entire croplands). But anticipated changes of agro-economic conditions, translating to altered crop share distributions, display stronger effects on soil C storage than climate change. Depending on future use of land expected to fall out of agricultural use in the future (~ 30 % of croplands area as “surplus” land), the basin either considerably looses soil C and the net annual C flux to the atmosphere increases (surplus used as black fallow) or the basin converts to a net sink of C (sequestering 0.44 106 tons CO2 yr-1 under extensified use as ley-arable) or reacts with decrease in source strength when using bioenergy crops. Bioenergy crops additionally offer a considerable potential for fossil fuel substitution (~37 PJ, 1015 J per year), whereas the basin wide use of harvest by-products for energy generation has to be seen critically although offering an annual energy potential of approximately 125 PJ. Harvest by-products play a central role in soil C reproduction and a percentage between 50 and 80 % should remain on the fields in order to maintain soil quality and fertility. The established modelling tool allows quantifying climate, land use and major land management impacts on soil C balance. New is that the SOM turnover description is embedded in an eco-hydrological river basin model, allowing an integrated consideration of water quantity, water quality, vegetation growth, agricultural productivity and soil carbon changes under different environmental conditions. The methodology and assessment presented here demonstrates the potential for integrated assessment of soil C dynamics alongside with other ecosystem services under global change impacts and provides information on the potentials of soils for climate change mitigation (soil C sequestration) and on their soil fertility status.
Natural and human induced environmental changes affect populations at different time scales. If they occur in a spatial heterogeneous way, they cause spatial variation in abundance. In this thesis I addressed three topics, all related to the question, how environmental changes influence population dynamics. In the first part, I analysed the effect of positive temporal autocorrelation in environmental noise on the extinction risk of a population, using a simple population model. The effect of autocorrelation depended on the magnitude of the effect of single catastrophic events of bad environmental conditions on a population. If a population was threatened by extinction only, when bad conditions occurred repeatedly, positive autocorrelation increased extinction risk. If a population could become extinct, even if bad conditions occurred only once, positive autocorrelation decreased extinction risk. These opposing effects could be explained by two features of an autocorrelated time series. On the one hand, positive autocorrelation increased the probability of series of bad environmental conditions, implying a negative effect on populations. On the other hand, aggregation of bad years also implied longer periods with relatively good conditions. Therefore, for a given time period, the overall probability of occurrence of at least one extremely bad year was reduced in autocorrelated noise. This can imply a positive effect on populations. The results could solve a contradiction in the literature, where opposing effects of autocorrelated noise were found in very similar population models. In the second part, I compared two approaches, which are commonly used for predicting effects of climate change on future abundance and distribution of species: a "space for time approach", where predictions are based on the geographic pattern of current abundance in relation to climate, and a "population modelling approach" which is based on correlations between demographic parameters and the inter-annual variation of climate. In this case study, I compared the two approaches for predicting the effect of a shift in mean precipitation on a population of the sociable weaver Philetairus socius, a common colonially living passerine bird of semiarid savannahs of southern Africa. In the space for time approach, I compared abundance and population structure of the sociable weaver in two areas with highly different mean annual precipitation. The analysis showed no difference between the two populations. This result, as well as the wide distribution range of the species, would lead to the prediction of no sensitive response of the species to a slight shift in mean precipitation. In contrast, the population modelling approach, based on a correlation between reproductive success and rainfall, predicted a sensitive response in most model types. The inconsistency of predictions was confirmed in a cross-validation between the two approaches. I concluded that the inconsistency was caused, because the two approaches reflect different time scales. On a short time scale, the population may respond sensitively to rainfall. However, on a long time scale, or in a regional comparison, the response may be compensated or buffered by a variety of mechanisms. These may include behavioural or life history adaptations, shifts in the interactions with other species, or differences in the physical environment. The study implies that understanding, how such mechanisms work, and at what time scale they would follow climate change, is a crucial precondition for predicting ecological consequences of climate change. In the third part of the thesis, I tested why colony sizes of the sociable weaver are highly variable. The high variation of colony sizes is surprising, as in studies on coloniality it is often assumed that an optimal colony size exists, in which individual bird fitness is maximized. Following this assumption, the pattern of bird dispersal should keep colony sizes near an optimum. However, I showed by analysing data on reproductive success and survival that for the sociable weaver fitness in relation to colony size did not follow an optimum curve. Instead, positive and negative effects of living in large colonies overlaid each other in a way that fitness was generally close to one, and density dependence was low. I showed in a population model, which included an evolutionary optimisation process of dispersal that this specific shape of the fitness function could lead to a dispersal strategy, where the variation of colony sizes was maintained.