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Institute
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A catalog of genetic loci associated with kidney function from analyses of a million individuals
(2019)
Wuttke, Matthias ; Li, Yong ; Li, Man ; Sieber, Karsten B. ; Feitosa, Mary F. ; Gorski, Mathias ; Tin, Adrienne ; Wang, Lihua ; Chu, Audrey Y. ; Hoppmann, Anselm ; Kirsten, Holger ; Giri, Ayush ; Chai, Jin-Fang ; Sveinbjornsson, Gardar ; Tayo, Bamidele O. ; Nutile, Teresa ; Fuchsberger, Christian ; Marten, Jonathan ; Cocca, Massimiliano ; Ghasemi, Sahar ; Xu, Yizhe ; Horn, Katrin ; Noce, Damia ; Van der Most, Peter J. ; Sedaghat, Sanaz ; Yu, Zhi ; Akiyama, Masato ; Afaq, Saima ; Ahluwalia, Tarunveer Singh ; Almgren, Peter ; Amin, Najaf ; Arnlov, Johan ; Bakker, Stephan J. L. ; Bansal, Nisha ; Baptista, Daniela ; Bergmann, Sven ; Biggs, Mary L. ; Biino, Ginevra ; Boehnke, Michael ; Boerwinkle, Eric ; Boissel, Mathilde ; Böttinger, Erwin ; Boutin, Thibaud S. ; Brenner, Hermann ; Brumat, Marco ; Burkhardt, Ralph ; Butterworth, Adam S. ; Campana, Eric ; Campbell, Archie ; Campbell, Harry ; Canouil, Mickael ; Carroll, Robert J. ; Catamo, Eulalia ; Chambers, John C. ; Chee, Miao-Ling ; Chee, Miao-Li ; Chen, Xu ; Cheng, Ching-Yu ; Cheng, Yurong ; Christensen, Kaare ; Cifkova, Renata ; Ciullo, Marina ; Concas, Maria Pina ; Cook, James P. ; Coresh, Josef ; Corre, Tanguy ; Sala, Cinzia Felicita ; Cusi, Daniele ; Danesh, John ; Daw, E. Warwick ; De Borst, Martin H. ; De Grandi, Alessandro ; De Mutsert, Renee ; De Vries, Aiko P. J. ; Degenhardt, Frauke ; Delgado, Graciela ; Demirkan, Ayse ; Di Angelantonio, Emanuele ; Dittrich, Katalin ; Divers, Jasmin ; Dorajoo, Rajkumar ; Eckardt, Kai-Uwe ; Ehret, Georg ; Elliott, Paul ; Endlich, Karlhans ; Evans, Michele K. ; Felix, Janine F. ; Foo, Valencia Hui Xian ; Franco, Oscar H. ; Franke, Andre ; Freedman, Barry I. ; Freitag-Wolf, Sandra ; Friedlander, Yechiel ; Froguel, Philippe ; Gansevoort, Ron T. ; Gao, He ; Gasparini, Paolo ; Gaziano, J. Michael ; Giedraitis, Vilmantas ; Gieger, Christian ; Girotto, Giorgia ; Giulianini, Franco ; Gogele, Martin ; Gordon, Scott D. ; Gudbjartsson, Daniel F. ; Gudnason, Vilmundur ; Haller, Toomas ; Hamet, Pavel ; Harris, Tamara B. ; Hartman, Catharina A. ; Hayward, Caroline ; Hellwege, Jacklyn N. ; Heng, Chew-Kiat ; Hicks, Andrew A. ; Hofer, Edith ; Huang, Wei ; Hutri-Kahonen, Nina ; Hwang, Shih-Jen ; Ikram, M. Arfan ; Indridason, Olafur S. ; Ingelsson, Erik ; Ising, Marcus ; Jaddoe, Vincent W. V. ; Jakobsdottir, Johanna ; Jonas, Jost B. ; Joshi, Peter K. ; Josyula, Navya Shilpa ; Jung, Bettina ; Kahonen, Mika ; Kamatani, Yoichiro ; Kammerer, Candace M. ; Kanai, Masahiro ; Kastarinen, Mika ; Kerr, Shona M. ; Khor, Chiea-Chuen ; Kiess, Wieland ; Kleber, Marcus E. ; Koenig, Wolfgang ; Kooner, Jaspal S. ; Korner, Antje ; Kovacs, Peter ; Kraja, Aldi T. ; Krajcoviechova, Alena ; Kramer, Holly ; Kramer, Bernhard K. ; Kronenberg, Florian ; Kubo, Michiaki ; Kuhnel, Brigitte ; Kuokkanen, Mikko ; Kuusisto, Johanna ; La Bianca, Martina ; Laakso, Markku ; Lange, Leslie A. ; Langefeld, Carl D. ; Lee, Jeannette Jen-Mai ; Lehne, Benjamin ; Lehtimaki, Terho ; Lieb, Wolfgang ; Lim, Su-Chi ; Lind, Lars ; Lindgren, Cecilia M. ; Liu, Jun ; Liu, Jianjun ; Loeffler, Markus ; Loos, Ruth J. F. ; Lucae, Susanne ; Lukas, Mary Ann ; Lyytikainen, Leo-Pekka ; Magi, Reedik ; Magnusson, Patrik K. E. ; Mahajan, Anubha ; Martin, Nicholas G. ; Martins, Jade ; Marz, Winfried ; Mascalzoni, Deborah ; Matsuda, Koichi ; Meisinger, Christa ; Meitinger, Thomas ; Melander, Olle ; Metspalu, Andres ; Mikaelsdottir, Evgenia K. ; Milaneschi, Yuri ; Miliku, Kozeta ; Mishra, Pashupati P. ; Program, V. A. Million Veteran ; Mohlke, Karen L. ; Mononen, Nina ; Montgomery, Grant W. ; Mook-Kanamori, Dennis O. ; Mychaleckyj, Josyf C. ; Nadkarni, Girish N. ; Nalls, Mike A. ; Nauck, Matthias ; Nikus, Kjell ; Ning, Boting ; Nolte, Ilja M. ; Noordam, Raymond ; Olafsson, Isleifur ; Oldehinkel, Albertine J. ; Orho-Melander, Marju ; Ouwehand, Willem H. ; Padmanabhan, Sandosh ; Palmer, Nicholette D. ; Palsson, Runolfur ; Penninx, Brenda W. J. H. ; Perls, Thomas ; Perola, Markus ; Pirastu, Mario ; Pirastu, Nicola ; Pistis, Giorgio ; Podgornaia, Anna I. ; Polasek, Ozren ; Ponte, Belen ; Porteous, David J. ; Poulain, Tanja ; Pramstaller, Peter P. ; Preuss, Michael H. ; Prins, Bram P. ; Province, Michael A. ; Rabelink, Ton J. ; Raffield, Laura M. ; Raitakari, Olli T. ; Reilly, Dermot F. ; Rettig, Rainer ; Rheinberger, Myriam ; Rice, Kenneth M. ; Ridker, Paul M. ; Rivadeneira, Fernando ; Rizzi, Federica ; Roberts, David J. ; Robino, Antonietta ; Rossing, Peter ; Rudan, Igor ; Rueedi, Rico ; Ruggiero, Daniela ; Ryan, Kathleen A. ; Saba, Yasaman ; Sabanayagam, Charumathi ; Salomaa, Veikko ; Salvi, Erika ; Saum, Kai-Uwe ; Schmidt, Helena ; Schmidt, Reinhold ; Ben Schottker ; Schulz, Christina-Alexandra ; Schupf, Nicole ; Shaffer, Christian M. ; Shi, Yuan ; Smith, Albert V. ; Smith, Blair H. ; Soranzo, Nicole ; Spracklen, Cassandra N. ; Strauch, Konstantin ; Stringham, Heather M. ; Stumvoll, Michael ; Svensson, Per O. ; Szymczak, Silke ; Tai, E-Shyong ; Tajuddin, Salman M. ; Tan, Nicholas Y. Q. ; Taylor, Kent D. ; Teren, Andrej ; Tham, Yih-Chung ; Thiery, Joachim ; Thio, Chris H. L. ; Thomsen, Hauke ; Thorleifsson, Gudmar ; Toniolo, Daniela ; Tonjes, Anke ; Tremblay, Johanne ; Tzoulaki, Ioanna ; Uitterlinden, Andre G. ; Vaccargiu, Simona ; Van Dam, Rob M. ; Van der Harst, Pim ; Van Duijn, Cornelia M. ; Edward, Digna R. Velez ; Verweij, Niek ; Vogelezang, Suzanne ; Volker, Uwe ; Vollenweider, Peter ; Waeber, Gerard ; Waldenberger, Melanie ; Wallentin, Lars ; Wang, Ya Xing ; Wang, Chaolong ; Waterworth, Dawn M. ; Bin Wei, Wen ; White, Harvey ; Whitfield, John B. ; Wild, Sarah H. ; Wilson, James F. ; Wojczynski, Mary K. ; Wong, Charlene ; Wong, Tien-Yin ; Xu, Liang ; Yang, Qiong ; Yasuda, Masayuki ; Yerges-Armstrong, Laura M. ; Zhang, Weihua ; Zonderman, Alan B. ; Rotter, Jerome I. ; Bochud, Murielle ; Psaty, Bruce M. ; Vitart, Veronique ; Wilson, James G. ; Dehghan, Abbas ; Parsa, Afshin ; Chasman, Daniel I. ; Ho, Kevin ; Morris, Andrew P. ; Devuyst, Olivier ; Akilesh, Shreeram ; Pendergrass, Sarah A. ; Sim, Xueling ; Boger, Carsten A. ; Okada, Yukinori ; Edwards, Todd L. ; Snieder, Harold ; Stefansson, Kari ; Hung, Adriana M. ; Heid, Iris M. ; Scholz, Markus ; Teumer, Alexander ; Kottgen, Anna ; Pattaro, Cristian
Chronic kidney disease (CKD) is responsible for a public health burden with multi-systemic complications. Through transancestry meta-analysis of genome-wide association studies of estimated glomerular filtration rate (eGFR) and independent replication (n = 1,046,070), we identified 264 associated loci (166 new). Of these,147 were likely to be relevant for kidney function on the basis of associations with the alternative kidney function marker blood urea nitrogen (n = 416,178). Pathway and enrichment analyses, including mouse models with renal phenotypes, support the kidney as the main target organ. A genetic risk score for lower eGFR was associated with clinically diagnosed CKD in 452,264 independent individuals. Colocalization analyses of associations with eGFR among 783,978 European-ancestry individuals and gene expression across 46 human tissues, including tubulo-interstitial and glomerular kidney compartments, identified 17 genes differentially expressed in kidney. Fine-mapping highlighted missense driver variants in 11 genes and kidney-specific regulatory variants. These results provide a comprehensive priority list of molecular targets for translational research.
Aarts, Alexander A. ; Anderson, Joanna E. ; Anderson, Christopher J. ; Attridge, Peter R. ; Attwood, Angela ; Axt, Jordan ; Babel, Molly ; Bahnik, Stepan ; Baranski, Erica ; Barnett-Cowan, Michael ; Bartmess, Elizabeth ; Beer, Jennifer ; Bell, Raoul ; Bentley, Heather ; Beyan, Leah ; Binion, Grace ; Borsboom, Denny ; Bosch, Annick ; Bosco, Frank A. ; Bowman, Sara D. ; Brandt, Mark J. ; Braswell, Erin ; Brohmer, Hilmar ; Brown, Benjamin T. ; Brown, Kristina ; Bruening, Jovita ; Calhoun-Sauls, Ann ; Callahan, Shannon P. ; Chagnon, Elizabeth ; Chandler, Jesse ; Chartier, Christopher R. ; Cheung, Felix ; Christopherson, Cody D. ; Cillessen, Linda ; Clay, Russ ; Cleary, Hayley ; Cloud, Mark D. ; Cohn, Michael ; Cohoon, Johanna ; Columbus, Simon ; Cordes, Andreas ; Costantini, Giulio ; Alvarez, Leslie D. Cramblet ; Cremata, Ed ; Crusius, Jan ; DeCoster, Jamie ; DeGaetano, Michelle A. ; Della Penna, Nicolas ; den Bezemer, Bobby ; Deserno, Marie K. ; Devitt, Olivia ; Dewitte, Laura ; Dobolyi, David G. ; Dodson, Geneva T. ; Donnellan, M. Brent ; Donohue, Ryan ; Dore, Rebecca A. ; Dorrough, Angela ; Dreber, Anna ; Dugas, Michelle ; Dunn, Elizabeth W. ; Easey, Kayleigh ; Eboigbe, Sylvia ; Eggleston, Casey ; Embley, Jo ; Epskamp, Sacha ; Errington, Timothy M. ; Estel, Vivien ; Farach, Frank J. ; Feather, Jenelle ; Fedor, Anna ; Fernandez-Castilla, Belen ; Fiedler, Susann ; Field, James G. ; Fitneva, Stanka A. ; Flagan, Taru ; Forest, Amanda L. ; Forsell, Eskil ; Foster, Joshua D. ; Frank, Michael C. ; Frazier, Rebecca S. ; Fuchs, Heather ; Gable, Philip ; Galak, Jeff ; Galliani, Elisa Maria ; Gampa, Anup ; Garcia, Sara ; Gazarian, Douglas ; Gilbert, Elizabeth ; Giner-Sorolla, Roger ; Glöckner, Andreas ; Göllner, Lars ; Goh, Jin X. ; Goldberg, Rebecca ; Goodbourn, Patrick T. ; Gordon-McKeon, Shauna ; Gorges, Bryan ; Gorges, Jessie ; Goss, Justin ; Graham, Jesse ; Grange, James A. ; Gray, Jeremy ; Hartgerink, Chris ; Hartshorne, Joshua ; Hasselman, Fred ; Hayes, Timothy ; Heikensten, Emma ; Henninger, Felix ; Hodsoll, John ; Holubar, Taylor ; Hoogendoorn, Gea ; Humphries, Denise J. ; Hung, Cathy O. -Y. ; Immelman, Nathali ; Irsik, Vanessa C. ; Jahn, Georg ; Jaekel, Frank ; Jekel, Marc ; Johannesson, Magnus ; Johnson, Larissa G. ; Johnson, David J. ; Johnson, Kate M. ; Johnston, William J. ; Jonas, Kai ; Joy-Gaba, Jennifer A. ; Kappes, Heather Barry ; Kelso, Kim ; Kidwell, Mallory C. ; Kim, Seung Kyung ; Kirkhart, Matthew ; Kleinberg, Bennett ; Knezevic, Goran ; Kolorz, Franziska Maria ; Kossakowski, Jolanda J. ; Krause, Robert Wilhelm ; Krijnen, Job ; Kuhlmann, Tim ; Kunkels, Yoram K. ; Kyc, Megan M. ; Lai, Calvin K. ; Laique, Aamir ; Lakens, Daniel ; Lane, Kristin A. ; Lassetter, Bethany ; Lazarevic, Ljiljana B. ; LeBel, Etienne P. ; Lee, Key Jung ; Lee, Minha ; Lemm, Kristi ; Levitan, Carmel A. ; Lewis, Melissa ; Lin, Lin ; Lin, Stephanie ; Lippold, Matthias ; Loureiro, Darren ; Luteijn, Ilse ; Mackinnon, Sean ; Mainard, Heather N. ; Marigold, Denise C. ; Martin, Daniel P. ; Martinez, Tylar ; Masicampo, E. J. ; Matacotta, Josh ; Mathur, Maya ; May, Michael ; Mechin, Nicole ; Mehta, Pranjal ; Meixner, Johannes ; Melinger, Alissa ; Miller, Jeremy K. ; Miller, Mallorie ; Moore, Katherine ; Möschl, Marcus ; Motyl, Matt ; Müller, Stephanie M. ; Munafo, Marcus ; Neijenhuijs, Koen I. ; Nervi, Taylor ; Nicolas, Gandalf ; Nilsonne, Gustav ; Nosek, Brian A. ; Nuijten, Michele B. ; Olsson, Catherine ; Osborne, Colleen ; Ostkamp, Lutz ; Pavel, Misha ; Penton-Voak, Ian S. ; Perna, Olivia ; Pernet, Cyril ; Perugini, Marco ; Pipitone, R. Nathan ; Pitts, Michael ; Plessow, Franziska ; Prenoveau, Jason M. ; Rahal, Rima-Maria ; Ratliff, Kate A. ; Reinhard, David ; Renkewitz, Frank ; Ricker, Ashley A. ; Rigney, Anastasia ; Rivers, Andrew M. ; Roebke, Mark ; Rutchick, Abraham M. ; Ryan, Robert S. ; Sahin, Onur ; Saide, Anondah ; Sandstrom, Gillian M. ; Santos, David ; Saxe, Rebecca ; Schlegelmilch, Rene ; Schmidt, Kathleen ; Scholz, Sabine ; Seibel, Larissa ; Selterman, Dylan Faulkner ; Shaki, Samuel ; Simpson, William B. ; Sinclair, H. Colleen ; Skorinko, Jeanine L. M. ; Slowik, Agnieszka ; Snyder, Joel S. ; Soderberg, Courtney ; Sonnleitner, Carina ; Spencer, Nick ; Spies, Jeffrey R. ; Steegen, Sara ; Stieger, Stefan ; Strohminger, Nina ; Sullivan, Gavin B. ; Talhelm, Thomas ; Tapia, Megan ; te Dorsthorst, Anniek ; Thomae, Manuela ; Thomas, Sarah L. ; Tio, Pia ; Traets, Frits ; Tsang, Steve ; Tuerlinckx, Francis ; Turchan, Paul ; Valasek, Milan ; Van Aert, Robbie ; van Assen, Marcel ; van Bork, Riet ; van de Ven, Mathijs ; van den Bergh, Don ; van der Hulst, Marije ; van Dooren, Roel ; van Doorn, Johnny ; van Renswoude, Daan R. ; van Rijn, Hedderik ; Vanpaemel, Wolf ; Echeverria, Alejandro Vasquez ; Vazquez, Melissa ; Velez, Natalia ; Vermue, Marieke ; Verschoor, Mark ; Vianello, Michelangelo ; Voracek, Martin ; Vuu, Gina ; Wagenmakers, Eric-Jan ; Weerdmeester, Joanneke ; Welsh, Ashlee ; Westgate, Erin C. ; Wissink, Joeri ; Wood, Michael ; Woods, Andy ; Wright, Emily ; Wu, Sining ; Zeelenberg, Marcel ; Zuni, Kellylynn
Reproducibility is a defining feature of science, but the extent to which it characterizes current research is unknown. We conducted replications of 100 experimental and correlational studies published in three psychology journals using high-powered designs and original materials when available. Replication effects were half the magnitude of original effects, representing a substantial decline. Ninety-seven percent of original studies had statistically significant results. Thirty-six percent of replications had statistically significant results; 47% of original effect sizes were in the 95% confidence interval of the replication effect size; 39% of effects were subjectively rated to have replicated the original result; and if no bias in original results is assumed, combining original and replication results left 68% with statistically significant effects. Correlational tests suggest that replication success was better predicted by the strength of original evidence than by characteristics of the original and replication teams.
Chipman, Ariel D. ; Ferrier, David E. K. ; Brena, Carlo ; Qu, Jiaxin ; Hughes, Daniel S. T. ; Schroeder, Reinhard ; Torres-Oliva, Montserrat ; Znassi, Nadia ; Jiang, Huaiyang ; Almeida, Francisca C. ; Alonso, Claudio R. ; Apostolou, Zivkos ; Aqrawi, Peshtewani ; Arthur, Wallace ; Barna, Jennifer C. J. ; Blankenburg, Kerstin P. ; Brites, Daniela ; Capella-Gutierrez, Salvador ; Coyle, Marcus ; Dearden, Peter K. ; Du Pasquier, Louis ; Duncan, Elizabeth J. ; Ebert, Dieter ; Eibner, Cornelius ; Erikson, Galina ; Evans, Peter D. ; Extavour, Cassandra G. ; Francisco, Liezl ; Gabaldon, Toni ; Gillis, William J. ; Goodwin-Horn, Elizabeth A. ; Green, Jack E. ; Griffiths-Jones, Sam ; Grimmelikhuijzen, Cornelis J. P. ; Gubbala, Sai ; Guigo, Roderic ; Han, Yi ; Hauser, Frank ; Havlak, Paul ; Hayden, Luke ; Helbing, Sophie ; Holder, Michael ; Hui, Jerome H. L. ; Hunn, Julia P. ; Hunnekuhl, Vera S. ; Jackson, LaRonda ; Javaid, Mehwish ; Jhangiani, Shalini N. ; Jiggins, Francis M. ; Jones, Tamsin E. ; Kaiser, Tobias S. ; Kalra, Divya ; Kenny, Nathan J. ; Korchina, Viktoriya ; Kovar, Christie L. ; Kraus, F. Bernhard ; Lapraz, Francois ; Lee, Sandra L. ; Lv, Jie ; Mandapat, Christigale ; Manning, Gerard ; Mariotti, Marco ; Mata, Robert ; Mathew, Tittu ; Neumann, Tobias ; Newsham, Irene ; Ngo, Dinh N. ; Ninova, Maria ; Okwuonu, Geoffrey ; Ongeri, Fiona ; Palmer, William J. ; Patil, Shobha ; Patraquim, Pedro ; Pham, Christopher ; Pu, Ling-Ling ; Putman, Nicholas H. ; Rabouille, Catherine ; Ramos, Olivia Mendivil ; Rhodes, Adelaide C. ; Robertson, Helen E. ; Robertson, Hugh M. ; Ronshaugen, Matthew ; Rozas, Julio ; Saada, Nehad ; Sanchez-Gracia, Alejandro ; Scherer, Steven E. ; Schurko, Andrew M. ; Siggens, Kenneth W. ; Simmons, DeNard ; Stief, Anna ; Stolle, Eckart ; Telford, Maximilian J. ; Tessmar-Raible, Kristin ; Thornton, Rebecca ; van der Zee, Maurijn ; von Haeseler, Arndt ; Williams, James M. ; Willis, Judith H. ; Wu, Yuanqing ; Zou, Xiaoyan ; Lawson, Daniel ; Muzny, Donna M. ; Worley, Kim C. ; Gibbs, Richard A. ; Akam, Michael ; Richards, Stephen
Myriapods (e. g., centipedes and millipedes) display a simple homonomous body plan relative to other arthropods. All members of the class are terrestrial, but they attained terrestriality independently of insects. Myriapoda is the only arthropod class not represented by a sequenced genome. We present an analysis of the genome of the centipede Strigamia maritima. It retains a compact genome that has undergone less gene loss and shuffling than previously sequenced arthropods, and many orthologues of genes conserved from the bilaterian ancestor that have been lost in insects. Our analysis locates many genes in conserved macro-synteny contexts, and many small-scale examples of gene clustering. We describe several examples where S. maritima shows different solutions from insects to similar problems. The insect olfactory receptor gene family is absent from S. maritima, and olfaction in air is likely effected by expansion of other receptor gene families. For some genes S. maritima has evolved paralogues to generate coding sequence diversity, where insects use alternate splicing. This is most striking for the Dscam gene, which in Drosophila generates more than 100,000 alternate splice forms, but in S. maritima is encoded by over 100 paralogues. We see an intriguing linkage between the absence of any known photosensory proteins in a blind organism and the additional absence of canonical circadian clock genes. The phylogenetic position of myriapods allows us to identify where in arthropod phylogeny several particular molecular mechanisms and traits emerged. For example, we conclude that juvenile hormone signalling evolved with the emergence of the exoskeleton in the arthropods and that RR-1 containing cuticle proteins evolved in the lineage leading to Mandibulata. We also identify when various gene expansions and losses occurred. The genome of S. maritima offers us a unique glimpse into the ancestral arthropod genome, while also displaying many adaptations to its specific life history.
Moreno, Marcelo Spegiorin ; Melnick, Daniel ; Rosenau, M. ; Báez, Juan Carlos ; Klotz, Jan ; Oncken, Onno ; Tassara, Andres ; Chen, J. ; Bataille, Klaus ; Bevis, M. ; Socquet, Anne ; Bolte, John ; Vigny, C. ; Brooks, B. ; Ryder, I. ; Grund, Volker ; Smalley, B. ; Carrizo, Daniel ; Bartsch, M. ; Hase, H.
The Maule earthquake of 27th February 2010 (M-w = 8.8) affected similar to 500 km of the Nazca-South America plate boundary in south-central Chile producing spectacular crustal deformation. Here, we present a detailed estimate of static coseismic surface offsets as measured by survey and continuous GPS, both in near- and far-field regions. Earthquake slip along the megathrust has been inferred from a Joint inversion of our new data together with published GPS, InSAR, and land-level changes data using Green's functions generated by a spherical finite-element model with realistic subduction zone geometry. The combination of the data sets provided a good resolution, indicating that most of the slip was well resolved. Coseismic slip was concentrated north of the epicenter with up to 16 m of slip, whereas to the south it reached over 10 m within two minor patches. A comparison of coseismic slip with the slip deficit accumulated since the last great earthquake in 1835 suggests that the 2010 event closed a mature seismic gap. Slip deficit distribution shows an apparent local overshoot that highlight cycle-to-cycle variability, which has to be taken into account when anticipating future events from interseismic observations. Rupture propagation was obviously not affected by bathymetric features of the incoming plate. Instead, splay faults in the upper plate seem to have limited rupture propagation in the updip and along-strike directions. Additionally, we found that along-strike gradients in slip are spatially correlated with geometrical inflections of the megathrust. Our study suggests that persistent tectonic features may control strain accumulation and release along subduction megathrusts.
Mrochen, Daniel M. ; Schulz, Daniel ; Fischer, Stefan ; Jeske, Kathrin ; El Gohary, Heba ; Reil, Daniela ; Imholt, Christian ; Truebe, Patricia ; Suchomel, Josef ; Tricaud, Emilie ; Jacob, Jens ; Heroldova, Marta ; Bröker, Barbara M. ; Strommenger, Birgit ; Walther, Birgit ; Ulrich, Rainer G. ; Holtfreter, Silva
Laboratory mice are the most commonly used animal model for Staphylococcus aureus infection studies. We have previously shown that laboratory mice from global vendors are frequently colonized with S. aureus. Laboratory mice originate from wild house mice. Hence, we investigated whether wild rodents, including house mice, as well as shrews are naturally colonized with S. aureus and whether S. aureus adapts to the wild animal host. 295 animals of ten different species were caught in different locations over four years (2012-2015) in Germany, France and the Czech Republic. 45 animals were positive for S. aureus (15.3%). Three animals were co-colonized with two different isolates, resulting in 48 S. aureus isolates in total. Positive animals were found in Germany and the Czech Republic in each studied year. The S. aureus isolates belonged to ten different spa types, which grouped into six lineages (clonal complex (CC) 49, CC88, CC130, CC1956, sequence type (ST) 890, ST3033). CC49 isolates were most abundant (17/48, 35.4%), followed by CC1956 (14/48, 29.2%) and ST890 (9/48, 18.8%). The wild animal isolates lacked certain properties that are common among human isolates, e.g., a phage-encoded immune evasion cluster, superantigen genes on mobile genetic elements and antibiotic resistance genes, which suggests long-term adaptation to the wild animal host. One CC130 isolate contained the mecC gene, implying wild rodents might be both reservoir and vector for methicillin-resistant. In conclusion, we demonstrated that wild rodents and shrews are naturally colonized with S. aureus, and that those S. aureus isolates show signs of host adaptation.
Actis, M. ; Agnetta, G. ; Aharonian, Felix A. ; Akhperjanian, A. G. ; Aleksic, J. ; Aliu, E. ; Allan, D. ; Allekotte, I. ; Antico, F. ; Antonelli, L. A. ; Antoranz, P. ; Aravantinos, A. ; Arlen, T. ; Arnaldi, H. ; Artmann, S. ; Asano, K. ; Asorey, H. G. ; Baehr, J. ; Bais, A. ; Baixeras, C. ; Bajtlik, S. ; Balis, D. ; Bamba, A. ; Barbier, C. ; Barcelo, M. ; Barnacka, Anna ; Barnstedt, Jürgen ; de Almeida, U. Barres ; Barrio, J. A. ; Basso, S. ; Bastieri, D. ; Bauer, C. ; Becerra Gonzalez, J. ; Becherini, Yvonne ; Bechtol, K. C. ; Becker, J. ; Beckmann, Volker ; Bednarek, W. ; Behera, B. ; Beilicke, M. ; Belluso, M. ; Benallou, M. ; Benbow, W. ; Berdugo, J. ; Berger, K. ; Bernardino, T. ; Bernlöhr, K. ; Biland, A. ; Billotta, S. ; Bird, T. ; Birsin, E. ; Bissaldi, E. ; Blake, S. ; Blanch Bigas, O. ; Bobkov, A. A. ; Bogacz, L. ; Bogdan, M. ; Boisson, Catherine ; Boix Gargallo, J. ; Bolmont, J. ; Bonanno, G. ; Bonardi, A. ; Bonev, T. ; Borkowski, Janett ; Botner, O. ; Bottani, A. ; Bourgeat, M. ; Boutonnet, C. ; Bouvier, A. ; Brau-Nogue, S. ; Braun, I. ; Bretz, T. ; Briggs, M. S. ; Brun, Pierre ; Brunetti, L. ; Buckley, H. ; Bugaev, V. ; Buehler, R. ; Bulik, Tomasz ; Busetto, G. ; Buson, S. ; Byrum, K. ; Cailles, M. ; Cameron, R. A. ; Canestrari, R. ; Cantu, S. ; Carmona, E. ; Carosi, A. ; Carr, John ; Carton, P. H. ; Casiraghi, M. ; Castarede, H. ; Catalano, O. ; Cavazzani, S. ; Cazaux, S. ; Cerruti, B. ; Cerruti, M. ; Chadwick, M. ; Chiang, J. ; Chikawa, M. ; Cieslar, M. ; Ciesielska, M. ; Cillis, A. N. ; Clerc, C. ; Colin, P. ; Colome, J. ; Compin, M. ; Conconi, P. ; Connaughton, V. ; Conrad, Jan ; Contreras, J. L. ; Coppi, P. ; Corlier, M. ; Corona, P. ; Corpace, O. ; Corti, D. ; Cortina, J. ; Costantini, H. ; Cotter, G. ; Courty, B. ; Couturier, S. ; Covino, S. ; Croston, J. ; Cusumano, G. ; Daniel, M. K. ; Dazzi, F. ; Deangelis, A. ; de Cea del Pozo, E. ; Dal Pino, E. M. de Gouveia ; de Jager, O. ; de la Calle Perez, I. ; De La Vega, G. ; De Lotto, B. ; de Naurois, M. ; Wilhelmi, E. de Ona ; de Souza, V. ; Decerprit, B. ; Deil, C. ; Delagnes, E. ; Deleglise, G. ; Delgado, C. ; Dettlaff, T. ; Di Paolo, A. ; Di Pierro, F. ; Diaz, C. ; Dick, J. ; Dickinson, H. ; Digel, S. W. ; Dimitrov, D. ; Disset, G. ; Djannati-Ataï, A. ; Doert, M. ; Domainko, W. ; Dorner, D. ; Doro, M. ; Dournaux, J. -L. ; Dravins, D. ; Drury, L. ; Dubois, F. ; Dubois, R. ; Dubus, G. ; Dufour, C. ; Durand, D. ; Dyks, J. ; Dyrda, M. ; Edy, E. ; Egberts, Kathrin ; Eleftheriadis, C. ; Elles, S. ; Emmanoulopoulos, D. ; Enomoto, R. ; Ernenwein, J. -P. ; Errando, M. ; Etchegoyen, A. ; Falcone, A. D. ; Farakos, K. ; Farnier, C. ; Federici, S. ; Feinstein, F. ; Ferenc, D. ; Fillin-Martino, E. ; Fink, D. ; Finley, C. ; Finley, J. P. ; Firpo, R. ; Florin, D. ; Foehr, C. ; Fokitis, E. ; Font, Ll. ; Fontaine, G. ; Fontana, A. ; Foerster, A. ; Fortson, L. ; Fouque, N. ; Fransson, C. ; Fraser, G. W. ; Fresnillo, L. ; Fruck, C. ; Fujita, Y. ; Fukazawa, Y. ; Funk, S. ; Gaebele, W. ; Gabici, S. ; Gadola, A. ; Galante, N. ; Gallant, Y. ; Garcia, B. ; Garcia Lopez, R. J. ; Garrido, D. ; Garrido, L. ; Gascon, D. ; Gasq, C. ; Gaug, M. ; Gaweda, J. ; Geffroy, N. ; Ghag, C. ; Ghedina, A. ; Ghigo, M. ; Gianakaki, E. ; Giarrusso, S. ; Giavitto, G. ; Giebels, B. ; Giro, E. ; Giubilato, P. ; Glanzman, T. ; Glicenstein, J. -F. ; Gochna, M. ; Golev, V. ; Gomez Berisso, M. ; Gonzalez, A. ; Gonzalez, F. ; Granena, F. ; Graciani, R. ; Granot, J. ; Gredig, R. ; Green, A. ; Greenshaw, T. ; Grimm, O. ; Grube, J. ; Grudzinska, M. ; Grygorczuk, J. ; Guarino, V. ; Guglielmi, L. ; Guilloux, F. ; Gunji, S. ; Gyuk, G. ; Hadasch, D. ; Haefner, D. ; Hagiwara, R. ; Hahn, J. ; Hallgren, A. ; Hara, S. ; Hardcastle, M. J. ; Hassan, T. ; Haubold, T. ; Hauser, M. ; Hayashida, M. ; Heller, R. ; Henri, G. ; Hermann, G. ; Herrero, A. ; Hinton, James Anthony ; Hoffmann, D. ; Hofmann, W. ; Hofverberg, P. ; Horns, D. ; Hrupec, D. ; Huan, H. ; Huber, B. ; Huet, J. -M. ; Hughes, G. ; Hultquist, K. ; Humensky, T. B. ; Huppert, J. -F. ; Ibarra, A. ; Illa, J. M. ; Ingjald, J. ; Inoue, S. ; Inoue, Y. ; Ioka, K. ; Jablonski, C. ; Jacholkowska, A. ; Janiak, M. ; Jean, P. ; Jensen, H. ; Jogler, T. ; Jung, I. ; Kaaret, P. ; Kabuki, S. ; Kakuwa, J. ; Kalkuhl, C. ; Kankanyan, R. ; Kapala, M. ; Karastergiou, A. ; Karczewski, M. ; Karkar, S. ; Karlsson, N. ; Kasperek, J. ; Katagiri, H. ; Katarzynski, K. ; Kawanaka, N. ; Kedziora, B. ; Kendziorra, E. ; Khelifi, B. ; Kieda, D. ; Kifune, T. ; Kihm, T. ; Klepser, S. ; Kluzniak, W. ; Knapp, J. ; Knappy, A. R. ; Kneiske, T. ; Knoedlseder, J. ; Koeck, F. ; Kodani, K. ; Kohri, K. ; Kokkotas, K. ; Komin, N. ; Konopelko, A. ; Kosack, K. ; Kossakowski, R. ; Kostka, P. ; Kotula, J. ; Kowal, G. ; Koziol, J. ; Kraehenbuehl, T. ; Krause, J. ; Krawczynski, H. ; Krennrich, F. ; Kretzschmann, A. ; Kubo, H. ; Kudryavtsev, V. A. ; Kushida, J. ; La Barbera, N. ; La Parola, V. ; La Rosa, G. ; Lopez, A. ; Lamanna, G. ; Laporte, P. ; Lavalley, C. ; Le Flour, T. ; Le Padellec, A. ; Lenain, J. -P. ; Lessio, L. ; Lieunard, B. ; Lindfors, E. ; Liolios, A. ; Lohse, T. ; Lombardi, S. ; Lopatin, A. ; Lorenz, E. ; Lubinski, P. ; Luz, O. ; Lyard, E. ; Maccarone, M. C. ; Maccarone, T. ; Maier, G. ; Majumdar, P. ; Maltezos, S. ; Malkiewicz, P. ; Mana, C. ; Manalaysay, A. ; Maneva, G. ; Mangano, A. ; Manigot, P. ; Marin, J. ; Mariotti, M. ; Markoff, S. ; Martinez, G. ; Martinez, M. ; Mastichiadis, A. ; Matsumoto, H. ; Mattiazzo, S. ; Mazin, D. ; McComb, T. J. L. ; McCubbin, N. ; McHardy, I. ; Medina, C. ; Melkumyan, D. ; Mendes, A. ; Mertsch, P. ; Meucci, M. ; Michalowski, J. ; Micolon, P. ; Mineo, T. ; Mirabal, N. ; Mirabel, F. ; Miranda, J. M. ; Mirzoyan, R. ; Mizuno, T. ; Moal, B. ; Moderski, R. ; Molinari, E. ; Monteiro, I. ; Moralejo, A. ; Morello, C. ; Mori, K. ; Motta, G. ; Mottez, F. ; Moulin, Emmanuel ; Mukherjee, R. ; Munar, P. ; Muraishi, H. ; Murase, K. ; Murphy, A. Stj. ; Nagataki, S. ; Naito, T. ; Nakamori, T. ; Nakayama, K. ; Naumann, C. L. ; Naumann, D. ; Nayman, P. ; Nedbal, D. ; Niedzwiecki, A. ; Niemiec, J. ; Nikolaidis, A. ; Nishijima, K. ; Nolan, S. J. ; Nowak, N. ; O'Brien, P. T. ; Ochoa, I. ; Ohira, Y. ; Ohishi, M. ; Ohka, H. ; Okumura, A. ; Olivetto, C. ; Ong, R. A. ; Orito, R. ; Orr, M. ; Osborne, J. P. ; Ostrowski, M. ; Otero, L. ; Otte, A. N. ; Ovcharov, E. ; Oya, I. ; Ozieblo, A. ; Paiano, S. ; Pallota, J. ; Panazol, J. L. ; Paneque, D. ; Panter, M. ; Paoletti, R. ; Papyan, G. ; Paredes, J. M. ; Pareschi, G. ; Parsons, R. D. ; Arribas, M. Paz ; Pedaletti, G. ; Pepato, A. ; Persic, M. ; Petrucci, P. O. ; Peyaud, B. ; Piechocki, W. ; Pita, S. ; Pivato, G. ; Platos, L. ; Platzer, R. ; Pogosyan, L. ; Pohl, Martin ; Pojmanski, G. ; Ponz, J. D. ; Potter, W. ; Prandini, E. ; Preece, R. ; Prokoph, H. ; Puehlhofer, G. ; Punch, M. ; Quel, E. ; Quirrenbach, A. ; Rajda, P. ; Rando, R. ; Rataj, M. ; Raue, M. ; Reimann, C. ; Reimann, O. ; Reimer, A. ; Reimer, O. ; Renaud, M. ; Renner, S. ; Reymond, J. -M. ; Rhode, W. ; Ribo, M. ; Ribordy, M. ; Rico, J. ; Rieger, F. ; Ringegni, P. ; Ripken, J. ; Ristori, P. ; Rivoire, S. ; Rob, L. ; Rodriguez, S. ; Roeser, U. ; Romano, Patrizia ; Romero, G. E. ; Rosier-Lees, S. ; Rovero, A. C. ; Roy, F. ; Royer, S. ; Rudak, B. ; Rulten, C. B. ; Ruppel, J. ; Russo, F. ; Ryde, F. ; Sacco, B. ; Saggion, A. ; Sahakian, V. ; Saito, K. ; Saito, T. ; Sakaki, N. ; Salazar, E. ; Salini, A. ; Sanchez, F. ; Sanchez Conde, M. A. ; Santangelo, Andrea ; Santos, E. M. ; Sanuy, A. ; Sapozhnikov, L. ; Sarkar, S. ; Scalzotto, V. ; Scapin, V. ; Scarcioffolo, M. ; Schanz, T. ; Schlenstedt, S. ; Schlickeiser, R. ; Schmidt, T. ; Schmoll, J. ; Schroedter, M. ; Schultz, C. ; Schultze, J. ; Schulz, A. ; Schwanke, U. ; Schwarzburg, S. ; Schweizer, T. ; Seiradakis, J. ; Selmane, S. ; Seweryn, K. ; Shayduk, M. ; Shellard, R. C. ; Shibata, T. ; Sikora, M. ; Silk, J. ; Sillanpaa, A. ; Sitarek, J. ; Skole, C. ; Smith, N. ; Sobczynska, D. ; Sofo Haro, M. ; Sol, H. ; Spanier, F. ; Spiga, D. ; Spyrou, S. ; Stamatescu, V. ; Stamerra, A. ; Starling, R. L. C. ; Stawarz, L. ; Steenkamp, R. ; Stegmann, Christian ; Steiner, S. ; Stergioulas, N. ; Sternberger, R. ; Stinzing, F. ; Stodulski, M. ; Straumann, U. ; Suarez, A. ; Suchenek, M. ; Sugawara, R. ; Sulanke, K. H. ; Sun, S. ; Supanitsky, A. D. ; Sutcliffe, P. ; Szanecki, M. ; Szepieniec, T. ; Szostek, A. ; Szymkowiak, A. ; Tagliaferri, G. ; Tajima, H. ; Takahashi, H. ; Takahashi, K. ; Takalo, L. ; Takami, H. ; Talbot, R. G. ; Tam, P. H. ; Tanaka, M. ; Tanimori, T. ; Tavani, M. ; Tavernet, J. -P. ; Tchernin, C. ; Tejedor, L. A. ; Telezhinsky, Igor O. ; Temnikov, P. ; Tenzer, C. ; Terada, Y. ; Terrier, R. ; Teshima, M. ; Testa, V. ; Tibaldo, L. ; Tibolla, O. ; Tluczykont, M. ; Peixoto, C. J. Todero ; Tokanai, F. ; Tokarz, M. ; Toma, K. ; Torres, D. F. ; Tosti, G. ; Totani, T. ; Toussenel, F. ; Vallania, P. ; Vallejo, G. ; van der Walt, J. ; van Eldik, C. ; Vandenbroucke, J. ; Vankov, H. ; Vasileiadis, G. ; Vassiliev, V. V. ; Vegas, I. ; Venter, L. ; Vercellone, S. ; Veyssiere, C. ; Vialle, J. P. ; Videla, M. ; Vincent, P. ; Vink, J. ; Vlahakis, N. ; Vlahos, L. ; Vogler, P. ; Vollhardt, A. ; Volpe, F. ; Von Gunten, H. P. ; Vorobiov, S. ; Wagner, S. ; Wagner, R. M. ; Wagner, B. ; Wakely, S. P. ; Walter, P. ; Walter, R. ; Warwick, R. ; Wawer, P. ; Wawrzaszek, R. ; Webb, N. ; Wegner, P. ; Weinstein, A. ; Weitzel, Q. ; Welsing, R. ; Wetteskind, H. ; White, R. ; Wierzcholska, A. ; Wilkinson, M. I. ; Williams, D. A. ; Winde, M. ; Wischnewski, R. ; Wisniewski, L. ; Wolczko, A. ; Wood, M. ; Xiong, Q. ; Yamamoto, T. ; Yamaoka, K. ; Yamazaki, R. ; Yanagita, S. ; Yoffo, B. ; Yonetani, M. ; Yoshida, A. ; Yoshida, T. ; Yoshikoshi, T. ; Zabalza, V. ; Zagdanski, A. ; Zajczyk, A. ; Zdziarski, A. ; Zech, Alraune ; Zietara, K. ; Ziolkowski, P. ; Zitelli, V. ; Zychowski, P.
Ground-based gamma-ray astronomy has had a major breakthrough with the impressive results obtained using systems of imaging atmospheric Cherenkov telescopes. Ground-based gamma-ray astronomy has a huge potential in astrophysics, particle physics and cosmology. CTA is an international initiative to build the next generation instrument, with a factor of 5-10 improvement in sensitivity in the 100 GeV-10 TeV range and the extension to energies well below 100 GeV and above 100 TeV. CTA will consist of two arrays (one in the north, one in the south) for full sky coverage and will be operated as open observatory. The design of CTA is based on currently available technology. This document reports on the status and presents the major design concepts of CTA.
The 2010 very high energy gamma-ray flare and 10 years ofmulti-wavelength oservations of M 87
(2012)
Abramowski, Attila ; Acero, F. ; Aharonian, Felix A. ; Akhperjanian, A. G. ; Anton, Gisela ; Balzer, Arnim ; Barnacka, Anna ; de Almeida, U. Barres ; Becherini, Yvonne ; Becker, J. ; Behera, B. ; Bernlöhr, K. ; Birsin, E. ; Biteau, Jonathan ; Bochow, A. ; Boisson, Catherine ; Bolmont, J. ; Bordas, Pol ; Brucker, J. ; Brun, Francois ; Brun, Pierre ; Bulik, Tomasz ; Buesching, I. ; Carrigan, Svenja ; Casanova, Sabrina ; Cerruti, M. ; Chadwick, Paula M. ; Charbonnier, A. ; Chaves, Ryan C. G. ; Cheesebrough, A. ; Clapson, A. C. ; Coignet, G. ; Cologna, Gabriele ; Conrad, Jan ; Dalton, M. ; Daniel, M. K. ; Davids, I. D. ; Degrange, B. ; Deil, C. ; Dickinson, H. J. ; Djannati-Ataï, A. ; Domainko, W. ; Drury, L. O'C. ; Dubus, G. ; Dutson, K. ; Dyks, J. ; Dyrda, M. ; Egberts, Kathrin ; Eger, P. ; Espigat, P. ; Fallon, L. ; Farnier, C. ; Fegan, S. ; Feinstein, F. ; Fernandes, M. V. ; Fiasson, A. ; Fontaine, G. ; Foerster, A. ; Fuessling, M. ; Gallant, Y. A. ; Gast, H. ; Gerard, L. ; Gerbig, D. ; Giebels, B. ; Glicenstein, J. F. ; Glueck, B. ; Goret, P. ; Goering, D. ; Haeffner, S. ; Hague, J. D. ; Hampf, D. ; Hauser, M. ; Heinz, S. ; Heinzelmann, G. ; Henri, G. ; Hermann, G. ; Hinton, James Anthony ; Hoffmann, A. ; Hofmann, W. ; Hofverberg, P. ; Holler, M. ; Horns, D. ; Jacholkowska, A. ; de Jager, O. C. ; Jahn, C. ; Jamrozy, M. ; Jung, I. ; Kastendieck, M. A. ; Katarzynski, K. ; Katz, U. ; Kaufmann, S. ; Keogh, D. ; Khangulyan, D. ; Khelifi, B. ; Klochkov, D. ; Kluzniak, W. ; Kneiske, T. ; Komin, Nu. ; Kosack, K. ; Kossakowski, R. ; Laffon, H. ; Lamanna, G. ; Lennarz, D. ; Lohse, T. ; Lopatin, A. ; Lu, C. -C. ; Marandon, V. ; Marcowith, Alexandre ; Masbou, J. ; Maurin, D. ; Maxted, N. ; Mayer, M. ; McComb, T. J. L. ; Medina, M. C. ; Mehault, J. ; Moderski, R. ; Moulin, Emmanuel ; Naumann, C. L. ; Naumann-Godo, M. ; de Naurois, M. ; Nedbal, D. ; Nekrassov, D. ; Nguyen, N. ; Nicholas, B. ; Niemiec, J. ; Nolan, S. J. ; Ohm, S. ; Wilhelmi, E. de Ona ; Opitz, B. ; Ostrowski, M. ; Oya, I. ; Panter, M. ; Arribas, M. Paz ; Pedaletti, G. ; Pelletier, G. ; Petrucci, P. -O. ; Pita, S. ; Puehlhofer, G. ; Punch, M. ; Quirrenbach, A. ; Raue, M. ; Rayner, S. M. ; Reimer, A. ; Reimer, O. ; Renaud, M. ; de los Reyes, R. ; Rieger, F. ; Ripken, J. ; Rob, L. ; Rosier-Lees, S. ; Rowell, G. ; Rudak, B. ; Rulten, C. B. ; Ruppel, J. ; Sahakian, V. ; Sanchez, David M. ; Santangelo, Andrea ; Schlickeiser, R. ; Schoeck, F. M. ; Schulz, A. ; Schwanke, U. ; Schwarzburg, S. ; Schwemmer, S. ; Sheidaei, F. ; Skilton, J. L. ; Sol, H. ; Spengler, G. ; Stawarz, L. ; Steenkamp, R. ; Stegmann, Christian ; Stinzing, F. ; Stycz, K. ; Sushch, Iurii ; Szostek, A. ; Tavernet, J. -P. ; Terrier, R. ; Tluczykont, M. ; Valerius, K. ; van Eldik, C. ; Vasileiadis, G. ; Venter, C. ; Vialle, J. P. ; Viana, A. ; Vincent, P. ; Voelk, H. J. ; Volpe, F. ; Vorobiov, S. ; Vorster, M. ; Wagner, S. J. ; Ward, M. ; White, R. ; Wierzcholska, A. ; Zacharias, M. ; Zajczyk, A. ; Zdziarski, A. A. ; Zech, Alraune ; Zechlin, H. -S. ; Aleksic, J. ; Antonelli, L. A. ; Antoranz, P. ; Backes, Michael ; Barrio, J. A. ; Bastieri, D. ; Becerra Gonzalez, J. ; Bednarek, W. ; Berdyugin, A. ; Berger, K. ; Bernardini, E. ; Biland, A. ; Blanch Bigas, O. ; Bock, R. K. ; Boller, A. ; Bonnoli, G. ; Tridon, D. Borla ; Braun, I. ; Bretz, T. ; Canellas, A. ; Carmona, E. ; Carosi, A. ; Colin, P. ; Colombo, E. ; Contreras, J. L. ; Cortina, J. ; Cossio, L. ; Covino, S. ; Dazzi, F. ; De Angelis, A. ; De Cea del Pozo, E. ; De Lotto, B. ; Delgado Mendez, C. ; Diago Ortega, A. ; Doert, M. ; Dominguez, A. ; Prester, Dijana Dominis ; Dorner, D. ; Doro, M. ; Elsaesser, D. ; Ferenc, D. ; Fonseca, M. V. ; Font, L. ; Fruck, C. ; Garcia Lopez, R. J. ; Garczarczyk, M. ; Garrido, D. ; Giavitto, G. ; Godinovic, N. ; Hadasch, D. ; Haefner, D. ; Herrero, A. ; Hildebrand, D. ; Hoehne-Moench, D. ; Hose, J. ; Hrupec, D. ; Huber, B. ; Jogler, T. ; Klepser, S. ; Kraehenbuehl, T. ; Krause, J. ; La Barbera, A. ; Lelas, D. ; Leonardo, E. ; Lindfors, E. ; Lombardi, S. ; Lopez, M. ; Lorenz, E. ; Makariev, M. ; Maneva, G. ; Mankuzhiyil, N. ; Mannheim, K. ; Maraschi, L. ; Mariotti, M. ; Martinez, M. ; Mazin, D. ; Meucci, M. ; Miranda, J. M. ; Mirzoyan, R. ; Miyamoto, H. ; Moldon, J. ; Moralejo, A. ; Munar, P. ; Nieto, D. ; Nilsson, K. ; Orito, R. ; Oya, I. ; Paneque, D. ; Paoletti, R. ; Pardo, S. ; Paredes, J. M. ; Partini, S. ; Pasanen, M. ; Pauss, F. ; Perez-Torres, M. A. ; Persic, M. ; Peruzzo, L. ; Pilia, M. ; Pochon, J. ; Prada, F. ; Moroni, P. G. Prada ; Prandini, E. ; Puljak, I. ; Reichardt, I. ; Reinthal, R. ; Rhode, W. ; Ribo, M. ; Rico, J. ; Ruegamer, S. ; Saggion, A. ; Saito, K. ; Saito, T. Y. ; Salvati, M. ; Satalecka, K. ; Scalzotto, V. ; Scapin, V. ; Schultz, C. ; Schweizer, T. ; Shayduk, M. ; Shore, S. N. ; Sillanpaa, A. ; Sitarek, J. ; Sobczynska, D. ; Spanier, F. ; Spiro, S. ; Stamerra, A. ; Steinke, B. ; Storz, J. ; Strah, N. ; Suric, T. ; Takalo, L. ; Takami, H. ; Tavecchio, F. ; Temnikov, P. ; Terzic, T. ; Tescaro, D. ; Teshima, M. ; Thom, M. ; Tibolla, O. ; Torres, D. F. ; Treves, A. ; Vankov, H. ; Vogler, P. ; Wagner, R. M. ; Weitzel, Q. ; Zabalza, V. ; Zandanel, F. ; Zanin, R. ; Arlen, T. ; Aune, T. ; Beilicke, M. ; Benbow, W. ; Bouvier, A. ; Bradbury, S. M. ; Buckley, J. H. ; Bugaev, V. ; Byrum, K. ; Cannon, A. ; Cesarini, A. ; Ciupik, L. ; Connolly, M. P. ; Cui, W. ; Dickherber, R. ; Duke, C. ; Errando, M. ; Falcone, A. ; Finley, J. P. ; Finnegan, G. ; Fortson, L. ; Furniss, A. ; Galante, N. ; Gall, D. ; Godambe, S. ; Griffin, S. ; Grube, J. ; Gyuk, G. ; Hanna, D. ; Holder, J. ; Huan, H. ; Hui, C. M. ; Kaaret, P. ; Karlsson, N. ; Kertzman, M. ; Khassen, Y. ; Kieda, D. ; Krawczynski, H. ; Krennrich, F. ; Lang, M. J. ; LeBohec, S. ; Maier, G. ; McArthur, S. ; McCann, A. ; Moriarty, P. ; Mukherjee, R. ; Nunez, P. D. ; Ong, R. A. ; Orr, M. ; Otte, A. N. ; Park, N. ; Perkins, J. S. ; Pichel, A. ; Pohl, Martin ; Prokoph, H. ; Ragan, K. ; Reyes, L. C. ; Reynolds, P. T. ; Roache, E. ; Rose, H. J. ; Ruppel, J. ; Schroedter, M. ; Sembroski, G. H. ; Sentuerk, G. D. ; Telezhinsky, Igor O. ; Tesic, G. ; Theiling, M. ; Thibadeau, S. ; Varlotta, A. ; Vassiliev, V. V. ; Vivier, M. ; Wakely, S. P. ; Weekes, T. C. ; Williams, D. A. ; Zitzer, B. ; de Almeida, U. Barres ; Cara, M. ; Casadio, C. ; Cheung, C. C. ; McConville, W. ; Davies, F. ; Doi, A. ; Giovannini, G. ; Giroletti, M. ; Hada, K. ; Hardee, P. ; Harris, D. E. ; Junor, W. ; Kino, M. ; Lee, N. P. ; Ly, C. ; Madrid, J. ; Massaro, F. ; Mundell, C. G. ; Nagai, H. ; Perlman, E. S. ; Steele, I. A. ; Walker, R. C. ; Wood, D. L.
The giant radio galaxy M 87 with its proximity (16 Mpc), famous jet, and very massive black hole ((3-6) x 10(9) M-circle dot) provides a unique opportunity to investigate the origin of very high energy (VHE; E > 100 GeV) gamma-ray emission generated in relativistic outflows and the surroundings of supermassive black holes. M 87 has been established as a VHE gamma-ray emitter since 2006. The VHE gamma-ray emission displays strong variability on timescales as short as a day. In this paper, results from a joint VHE monitoring campaign on M 87 by the MAGIC and VERITAS instruments in 2010 are reported. During the campaign, a flare at VHE was detected triggering further observations at VHE (H.E.S.S.), X-rays (Chandra), and radio (43 GHz Very Long Baseline Array, VLBA). The excellent sampling of the VHE gamma-ray light curve enables one to derive a precise temporal characterization of the flare: the single, isolated flare is well described by a two-sided exponential function with significantly different flux rise and decay times of tau(rise)(d) = (1.69 +/- 0.30) days and tau(decay)(d) = (0.611 +/- 0.080) days, respectively. While the overall variability pattern of the 2010 flare appears somewhat different from that of previous VHE flares in 2005 and 2008, they share very similar timescales (similar to day), peak fluxes (Phi(>0.35 TeV) similar or equal to (1-3) x 10(-11) photons cm(-2) s(-1)), and VHE spectra. VLBA radio observations of 43 GHz of the inner jet regions indicate no enhanced flux in 2010 in contrast to observations in 2008, where an increase of the radio flux of the innermost core regions coincided with a VHE flare. On the other hand, Chandra X-ray observations taken similar to 3 days after the peak of the VHE gamma-ray emission reveal an enhanced flux from the core (flux increased by factor similar to 2; variability timescale <2 days). The long-term (2001-2010) multi-wavelength (MWL) light curve of M 87, spanning from radio to VHE and including data from Hubble Space Telescope, Liverpool Telescope, Very Large Array, and European VLBI Network, is used to further investigate the origin of the VHE gamma-ray emission. No unique, common MWL signature of the three VHE flares has been identified. In the outer kiloparsec jet region, in particular in HST-1, no enhanced MWL activity was detected in 2008 and 2010, disfavoring it as the origin of the VHE flares during these years. Shortly after two of the three flares (2008 and 2010), the X-ray core was observed to be at a higher flux level than its characteristic range (determined from more than 60 monitoring observations: 2002-2009). In 2005, the strong flux dominance of HST-1 could have suppressed the detection of such a feature. Published models for VHE gamma-ray emission from M 87 are reviewed in the light of the new data.
Prospects for Cherenkov Telescope Array Observations of the Young Supernova Remnant RX J1713.7-3946
(2017)
Acero, F. ; Aloisio, R. ; Amans, J. ; Amato, Elena ; Antonelli, L. A. ; Aramo, C. ; Armstrong, T. ; Arqueros, F. ; Asano, Katsuaki ; Ashley, M. ; Backes, M. ; Balazs, C. ; Balzer, A. ; Bamba, Aya ; Barkov, Maxim ; Barrio, J. A. ; Benbow, Wystan ; Bernloehr, K. ; Beshley, V. ; Bigongiari, C. ; Biland, A. ; Bilinsky, A. ; Bissaldi, Elisabetta ; Biteau, J. ; Blanch, O. ; Blasi, P. ; Blazek, J. ; Boisson, C. ; Bonanno, G. ; Bonardi, A. ; Bonavolonta, C. ; Bonnoli, G. ; Braiding, C. ; Brau-Nogue, S. ; Bregeon, J. ; Brown, A. M. ; Bugaev, V. ; Bulgarelli, A. ; Bulik, T. ; Burton, Michael ; Burtovoi, A. ; Busetto, G. ; Bottcher, M. ; Cameron, R. ; Capalbi, M. ; Caproni, Anderson ; Caraveo, P. ; Carosi, R. ; Cascone, E. ; Cerruti, M. ; Chaty, Sylvain ; Chen, A. ; Chen, X. ; Chernyakova, M. ; Chikawa, M. ; Chudoba, J. ; Cohen-Tanugi, J. ; Colafrancesco, S. ; Conforti, V. ; Contreras, J. L. ; Costa, A. ; Cotter, G. ; Covino, Stefano ; Covone, G. ; Cumani, P. ; Cusumano, G. ; Daniel, M. ; Dazzi, F. ; De Angelis, A. ; De Cesare, G. ; De Franco, A. ; De Frondat, F. ; Dal Pino, E. M. de Gouveia ; De Lisio, C. ; Lopez, R. de los Reyes ; De Lotto, B. ; de Naurois, M. ; De Palma, F. ; Del Santo, M. ; Delgado, C. ; della Volpe, D. ; Di Girolamo, T. ; Di Giulio, C. ; Di Pierro, F. ; Di Venere, L. ; Doro, M. ; Dournaux, J. ; Dumas, D. ; Dwarkadas, Vikram V. ; Diaz, C. ; Ebr, J. ; Egberts, Kathrin ; Einecke, S. ; Elsaesser, D. ; Eschbach, S. ; Falceta-Goncalves, D. ; Fasola, G. ; Fedorova, E. ; Fernandez-Barral, A. ; Ferrand, Gilles ; Fesquet, M. ; Fiandrini, E. ; Fiasson, A. ; Filipovic, Miroslav D. ; Fioretti, V. ; Font, L. ; Fontaine, Gilles ; Franco, F. J. ; Freixas Coromina, L. ; Fujita, Yutaka ; Fukui, Y. ; Funk, S. ; Forster, A. ; Gadola, A. ; Lopez, R. Garcia ; Garczarczyk, M. ; Giglietto, N. ; Giordano, F. ; Giuliani, A. ; Glicenstein, J. ; Gnatyk, R. ; Goldoni, P. ; Grabarczyk, T. ; Graciani, R. ; Graham, J. ; Grandi, P. ; Granot, Jonathan ; Green, A. J. ; Griffiths, S. ; Gunji, S. ; Hakobyan, H. ; Hara, S. ; Hassan, T. ; Hayashida, M. ; Heller, M. ; Helo, J. C. ; Hinton, J. ; Hnatyk, B. ; Huet, J. ; Huetten, M. ; Humensky, T. B. ; Hussein, M. ; Horandel, J. ; Ikeno, Y. ; Inada, T. ; Inome, Y. ; Inoue, S. ; Inoue, T. ; Inoue, Y. ; Ioka, K. ; Iori, Maurizio ; Jacquemier, J. ; Janecek, P. ; Jankowsky, D. ; Jung, I. ; Kaaret, P. ; Katagiri, H. ; Kimeswenger, S. ; Kimura, Shigeo S. ; Knodlseder, J. ; Koch, B. ; Kocot, J. ; Kohri, K. ; Komin, N. ; Konno, Y. ; Kosack, K. ; Koyama, S. ; Kraus, Michaela ; Kubo, Hidetoshi ; Mezek, G. Kukec ; Kushida, J. ; La Palombara, N. ; Lalik, K. ; Lamanna, G. ; Landt, H. ; Lapington, J. ; Laporte, P. ; Lee, S. ; Lees, J. ; Lefaucheur, J. ; Lenain, J. -P. ; Leto, Giuseppe ; Lindfors, E. ; Lohse, T. ; Lombardi, S. ; Longo, F. ; Lopez, M. ; Lucarelli, F. ; Luque-Escamilla, Pedro Luis ; Lopez-Coto, R. ; Maccarone, M. C. ; Maier, G. ; Malaguti, G. ; Mandat, D. ; Maneva, G. ; Mangano, S. ; Marcowith, Alexandre ; Marti, J. ; Martinez, M. ; Martinez, G. ; Masuda, S. ; Maurin, G. ; Maxted, N. ; Melioli, Claudio ; Mineo, T. ; Mirabal, N. ; Mizuno, T. ; Moderski, R. ; Mohammed, M. ; Montaruli, T. ; Moralejo, A. ; Mori, K. ; Morlino, G. ; Morselli, A. ; Moulin, Emmanuel ; Mukherjee, R. ; Mundell, C. ; Muraishi, H. ; Murase, Kohta ; Nagataki, Shigehiro ; Nagayoshi, T. ; Naito, T. ; Nakajima, D. ; Nakamori, T. ; Nemmen, R. ; Niemiec, Jacek ; Nieto, D. ; Nievas-Rosillo, M. ; Nikolajuk, M. ; Nishijima, K. ; Noda, K. ; Nogues, L. ; Nosek, D. ; Novosyadlyj, B. ; Nozaki, S. ; Ohira, Yutaka ; Ohishi, M. ; Ohm, S. ; Okumura, A. ; Ong, R. A. ; Orito, R. ; Orlati, A. ; Ostrowski, M. ; Oya, I. ; Padovani, Marco ; Palacio, J. ; Palatka, M. ; Paredes, Josep M. ; Pavy, S. ; Persic, M. ; Petrucci, P. ; Petruk, Oleh ; Pisarski, A. ; Pohl, Martin ; Porcelli, A. ; Prandini, E. ; Prast, J. ; Principe, G. ; Prouza, M. ; Pueschel, Elisa ; Puelhofer, G. ; Quirrenbach, A. ; Rameez, M. ; Reimer, O. ; Renaud, M. ; Ribo, M. ; Rico, J. ; Rizi, V. ; Rodriguez, J. ; Fernandez, G. Rodriguez ; Rodriguez Vazquez, J. J. ; Romano, Patrizia ; Romeo, G. ; Rosado, J. ; Rousselle, J. ; Rowell, G. ; Rudak, B. ; Sadeh, I. ; Safi-Harb, S. ; Saito, T. ; Sakaki, N. ; Sanchez, D. ; Sangiorgi, P. ; Sano, H. ; Santander, M. ; Sarkar, S. ; Sawada, M. ; Schioppa, E. J. ; Schoorlemmer, H. ; Schovanek, P. ; Schussler, F. ; Sergijenko, O. ; Servillat, M. ; Shalchi, A. ; Shellard, R. C. ; Siejkowski, H. ; Sillanpaa, A. ; Simone, D. ; Sliusar, V. ; Sol, H. ; Stanic, S. ; Starling, R. ; Stawarz, L. ; Stefanik, S. ; Stephan, M. ; Stolarczyk, T. ; Szanecki, M. ; Szepieniec, T. ; Tagliaferri, G. ; Tajima, H. ; Takahashi, M. ; Takeda, J. ; Tanaka, M. ; Tanaka, S. ; Tejedor, L. A. ; Telezhinsky, Igor O. ; Temnikov, P. ; Terada, Y. ; Tescaro, D. ; Teshima, M. ; Testa, V. ; Thoudam, S. ; Tokanai, F. ; Torres, D. F. ; Torresi, E. ; Tosti, G. ; Townsley, C. ; Travnicek, P. ; Trichard, C. ; Trifoglio, M. ; Tsujimoto, S. ; Vagelli, V. ; Vallania, P. ; Valore, L. ; van Driel, W. ; van Eldik, C. ; Vandenbroucke, Justin ; Vassiliev, V. ; Vecchi, M. ; Vercellone, Stefano ; Vergani, S. ; Vigorito, C. ; Vorobiov, S. ; Vrastil, M. ; Vazquez Acosta, M. L. ; Wagner, S. J. ; Wagner, R. ; Wakely, S. P. ; Walter, R. ; Ward, J. E. ; Watson, J. J. ; Weinstein, A. ; White, M. ; White, R. ; Wierzcholska, A. ; Wilcox, P. ; Williams, D. A. ; Wischnewski, R. ; Wojcik, P. ; Yamamoto, T. ; Yamamoto, H. ; Yamazaki, Ryo ; Yanagita, S. ; Yang, L. ; Yoshida, T. ; Yoshida, M. ; Yoshiike, S. ; Yoshikoshi, T. ; Zacharias, M. ; Zampieri, L. ; Zanin, R. ; Zavrtanik, M. ; Zavrtanik, D. ; Zdziarski, A. ; Zech, Alraune ; Zechlin, Hannes ; Zhdanov, V. ; Ziegler, A. ; Zorn, J.
We perform simulations for future Cherenkov Telescope Array (CTA) observations of RX J1713.7-3946, a young supernova remnant (SNR) and one of the brightest sources ever discovered in very high energy (VHE) gamma rays. Special attention is paid to exploring possible spatial (anti) correlations of gamma rays with emission at other wavelengths, in particular X-rays and CO/H I emission. We present a series of simulated images of RX J1713.7-3946 for CTA based on a set of observationally motivated models for the gamma-ray emission. In these models, VHE gamma rays produced by high-energy electrons are assumed to trace the nonthermal X-ray emission observed by XMM-Newton, whereas those originating from relativistic protons delineate the local gas distributions. The local atomic and molecular gas distributions are deduced by the NANTEN team from CO and H I observations. Our primary goal is to show how one can distinguish the emission mechanism(s) of the gamma rays (i.e., hadronic versus leptonic, or a mixture of the two) through information provided by their spatial distribution, spectra, and time variation. This work is the first attempt to quantitatively evaluate the capabilities of CTA to achieve various proposed scientific goals by observing this important cosmic particle accelerator.
Ahnen, M. L. ; Ansoldi, S. ; Antonelli, L. A. ; Arcaro, C. ; Babic, A. ; Banerjee, B. ; Bangale, P. ; Barres de Almeida, U. ; Barrio, J. A. ; Gonzalez, J. Becerra ; Bednarek, W. ; Bernardini, E. ; Berti, A. ; Bhattacharyya, W. ; Blanch, O. ; Bonnoli, G. ; Carosi, R. ; Carosi, A. ; Chatterjee, A. ; Colak, S. M. ; Colin, P. ; Colombo, E. ; Contreras, J. L. ; Cortina, J. ; Covino, S. ; Cumani, P. ; Da Vela, P. ; Dazzi, F. ; De Angelis, A. ; De Lotto, B. ; Delfino, M. ; Delgado, Jose Miguel Martins ; Di Pierro, F. ; Doert, M. ; Dominguez, A. ; Prester, D. Dominis ; Doro, M. ; Glawion, D. Eisenacher ; Engelkemeier, M. ; Ramazani, V. Fallah ; Fernandez-Barral, A. ; Fidalgo, D. ; Fonseca, M. V. ; Font, L. ; Fruck, C. ; Galindo, D. ; Lopez, R. J. Garcia ; Garczarczyk, M. ; Gaug, M. ; Giammaria, P. ; Godinovic, N. ; Gora, D. ; Guberman, D. ; Hadasch, D. ; Hahn, A. ; Hassan, T. ; Hayashida, M. ; Herrera, J. ; Hose, J. ; Hrupec, D. ; Ishio, K. ; Konno, Y. ; Kubo, H. ; Kushida, J. ; Kuvezdic, D. ; Lelas, D. ; Lindfors, E. ; Lombardi, S. ; Longo, F. ; Lopez, M. ; Maggio, C. ; Majumdar, P. ; Makariev, M. ; Maneva, G. ; Manganaro, M. ; Maraschi, L. ; Mariotti, M. ; Martinez, M. ; Mazin, D. ; Menzel, U. ; Minev, M. ; Miranda, J. M. ; Mirzoyan, R. ; Moralejo, A. ; Moreno, V. ; Moretti, E. ; Nagayoshi, T. ; Neustroev, V. ; Niedzwiecki, A. ; Nievas Rosillo, M. ; Nigro, C. ; Nilsson, K. ; Ninci, D. ; Nishijima, K. ; Noda, K. ; Nogues, L. ; Paiano, S. ; Palacio, J. ; Paneque, D. ; Paoletti, R. ; Paredes, J. M. ; Pedaletti, G. ; Peresano, M. ; Perri, L. ; Persic, M. ; Moroni, P. G. Prada ; Prandini, E. ; Puljak, I. ; Garcia, J. R. ; Reichardt, I. ; Ribo, M. ; Rico, J. ; Righi, C. ; Rugliancich, A. ; Saito, T. ; Satalecka, K. ; Schroeder, S. ; Schweizer, T. ; Shore, S. N. ; Sitarek, J. ; Snidaric, I. ; Sobczynska, D. ; Stamerra, A. ; Strzys, M. ; Suric, T. ; Takalo, L. ; Tavecchio, F. ; Temnikov, P. ; Terzic, T. ; Teshima, M. ; Torres-Alba, N. ; Treves, A. ; Tsujimoto, S. ; Vanzo, G. ; Vazquez Acosta, M. ; Vovk, I. ; Ward, J. E. ; Will, M. ; Zaric, D. ; Arbet-Engels, A. ; Baack, D. ; Balbo, M. ; Biland, A. ; Blank, M. ; Bretz, T. ; Bruegge, K. ; Bulinski, M. ; Buss, J. ; Dmytriiev, A. ; Dorner, D. ; Einecke, S. ; Elsaesser, D. ; Herbst, T. ; Hildebrand, D. ; Kortmann, L. ; Linhoff, L. ; Mahlke, M. ; Mannheim, K. ; Mueller, S. A. ; Neise, D. ; Neronov, A. ; Noethe, M. ; Oberkirch, J. ; Paravac, A. ; Rhode, W. ; Schleicher, B. ; Schulz, F. ; Sedlaczek, K. ; Shukla, A. ; Sliusar, V. ; Walter, R. ; Archer, A. ; Benbow, W. ; Bird, R. ; Brose, Robert ; Buckley, J. H. ; Bugaev, V. ; Christiansen, J. L. ; Cui, W. ; Daniel, M. K. ; Falcone, A. ; Feng, Q. ; Finley, J. P. ; Gillanders, G. H. ; Gueta, O. ; Hanna, D. ; Hervet, O. ; Holder, J. ; Hughes, G. ; Huetten, M. ; Humensky, T. B. ; Johnson, C. A. ; Kaaret, P. ; Kar, P. ; Kelley-Hoskins, N. ; Kertzman, M. ; Kieda, D. ; Krause, M. ; Krennrich, F. ; Kumar, S. ; Lang, M. J. ; Lin, T. T. Y. ; Maier, G. ; McArthur, S. ; Moriarty, P. ; Mukherjee, R. ; Ong, R. A. ; Otte, A. N. ; Park, N. ; Petrashyk, A. ; Pichel, A. ; Pohl, Martin ; Quinn, J. ; Ragan, K. ; Reynolds, P. T. ; Richards, G. T. ; Roache, E. ; Rovero, A. C. ; Rulten, C. ; Sadeh, I. ; Santander, M. ; Sembroski, G. H. ; Shahinyan, K. ; Sushch, Iurii ; Tyler, J. ; Wakely, S. P. ; Weinstein, A. ; Wells, R. M. ; Wilcox, P. ; Wilhel, A. ; Williams, D. A. ; Williamson, T. J. ; Zitzer, B. ; Perri, M. ; Verrecchia, F. ; Leto, C. ; Villata, M. ; Raiteri, C. M. ; Jorstad, S. G. ; Larionov, V. M. ; Blinov, D. A. ; Grishina, T. S. ; Kopatskaya, E. N. ; Larionova, E. G. ; Nikiforova, A. A. ; Morozova, D. A. ; Troitskaya, Yu. V. ; Troitsky, I. S. ; Kurtanidze, O. M. ; Nikolashvili, M. G. ; Kurtanidze, S. O. ; Kimeridze, G. N. ; Chigladze, R. A. ; Strigachev, A. ; Sadun, A. C.
Aims. We aim to characterize the multiwavelength emission from Markarian 501 (Mrk 501), quantify the energy-dependent variability, study the potential multiband correlations, and describe the temporal evolution of the broadband emission within leptonic theoretical scenarios. Methods. We organized a multiwavelength campaign to take place between March and July of 2012. Excellent temporal coverage was obtained with more than 25 instruments, including the MAGIC, FACT and VERITAS Cherenkov telescopes, the instruments on board the Swift and Fermi spacecraft, and the telescopes operated by the GASP-WEBT collaboration. Results. Mrk 501 showed a very high energy (VHE) gamma-ray flux above 0.2 TeV of similar to 0.5 times the Crab Nebula flux (CU) for most of the campaign. The highest activity occurred on 2012 June 9, when the VHE flux was similar to 3 CU, and the peak of the high-energy spectral component was found to be at similar to 2 TeV. Both the X-ray and VHE gamma-ray spectral slopes were measured to be extremely hard, with spectral indices <2 during most of the observing campaign, regardless of the X-ray and VHE flux. This study reports the hardest Mrk 501 VHE spectra measured to date. The fractional variability was found to increase with energy, with the highest variability occurring at VHE. Using the complete data set, we found correlation between the X-ray and VHE bands; however, if the June 9 flare is excluded, the correlation disappears (significance <3 sigma) despite the existence of substantial variability in the X-ray and VHE bands throughout the campaign. Conclusions. The unprecedentedly hard X-ray and VHE spectra measured imply that their low- and high-energy components peaked above 5 keV and 0.5 TeV, respectively, during a large fraction of the observing campaign, and hence that Mrk 501 behaved like an extreme high-frequency-peaked blazar (EHBL) throughout the 2012 observing season. This suggests that being an EHBL may not be a permanent characteristic of a blazar, but rather a state which may change over time. The data set acquired shows that the broadband spectral energy distribution (SED) of Mrk 501, and its transient evolution, is very complex, requiring, within the framework of synchrotron self-Compton (SSC) models, various emission regions for a satisfactory description. Nevertheless the one-zone SSC scenario can successfully describe the segments of the SED where most energy is emitted, with a significant correlation between the electron energy density and the VHE gamma-ray activity, suggesting that most of the variability may be explained by the injection of high-energy electrons. The one-zone SSC scenario used reproduces the behavior seen between the measured X-ray and VHE gamma-ray fluxes, and predicts that the correlation becomes stronger with increasing energy of the X-rays.
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