TY - JOUR A1 - Ashastina, Kseniia A1 - Kuzmina, Svetlana A1 - Rudaya, Natalia A1 - Troeva, Elena I. A1 - Schoch, Werner H. A1 - Roemermann, Christine A1 - Reinecke, Jennifer A1 - Otte, Volker A1 - Savvinov, Grigoriy A1 - Wesche, Karsten A1 - Kienast, Frank T1 - Woodlands and steppes BT - Pleistocene vegetation in Yakutia's most continental part recorded in the Batagay permafrost sequence JF - Quaternary science reviews : the international multidisciplinary research and review journal N2 - Based on fossil organism remains including plant macrofossils, charcoal, pollen, and invertebrates preserved in syngenetic deposits of the Batagay permafrost sequence in the Siberian Yana Highlands, we reconstructed the environmental history during marine isotope stages (MIS) 6 to 2. Two fossil assemblages, exceptionally rich in plant remains, allowed for a detailed description of the palaeo-vegetation during two climate extremes of the Late Pleistocene, the onset of the last glacial maximum (LGM) and the last interglacial. In addition, altogether 41 assemblages were used to outline the vegetation history since the penultimate cold stage of MIS 6. Accordingly, meadow steppes analogue to modern communities of the phytosociological order Festucetalia lenensis formed the primary vegetation during the Saalian and Weichselian cold stages. Cold-resistant tundra-steppe communities (Carici rupestris-Kobresietea bellardii) as they occur above the treeline today were, in contrast to more northern locations, mostly lacking. During the last interglacial, open coniferous woodland similar to modern larch taiga was the primary vegetation at the site. Abundant charcoal indicates wildfire events during the last interglacial. Zoogenic disturbances of the local vegetation were indicated by the presence of ruderal plants, especially by abundant Urtica dioica, suggesting that the area was an interglacial refugium for large herbivores. Meadow steppes, which formed the primary vegetation during cold stages and provided potentially suitable pastures for herbivores, were a significant constituent of the plant cover in the Yana Highlands also under the full warm stage conditions of the last interglacial. Consequently, meadow steppes occurred in the Yana Highlands during the entire investigated timespan from MIS 6 to MIS 2 documenting a remarkable environmental stability. Thus, the proportion of meadow steppe vegetation merely shifted in response to the respectively prevailing climatic conditions. Their persistence indicates low precipitation and a relatively warm growing season throughout and beyond the late Pleistocene. The studied fossil record also proves that modern steppe occurrences in the Yana Highlands did not establish as late as in the Holocene but instead are relicts of a formerly continuous steppe belt extending from Central Siberia to Northeast Yakutia during the Pleistocene. The persistence of plants and invertebrates characteristic of meadow steppe vegetation in interior Yakutia throughout the late Quaternary indicates climatic continuity and documents the suitability of this region as a refugium also for other organisms of the Pleistocene mammoth steppe including the iconic large herbivores. (C)2018 Elsevier Ltd. All rights reserved. KW - Palaeo-vegetation KW - Plant macrofossils KW - Invertebrates KW - Modern analogues KW - Pollen KW - Ground squirrel nest KW - Last cold stage KW - Eemian KW - Beringia Y1 - 2018 U6 - https://doi.org/10.1016/j.quascirev.2018.07.032 SN - 0277-3791 VL - 196 SP - 38 EP - 61 PB - Elsevier CY - Oxford ER - TY - JOUR A1 - Biskaborn, Boris A1 - Subetto, D. A. A1 - Savelieva, L. A. A1 - Vakhrameeva, P. S. A1 - Hansche, A. A1 - Herzschuh, Ulrike A1 - Klemm, J. A1 - Heinecke, L. A1 - Pestryakova, Luidmila Agafyevna A1 - Meyer, H. A1 - Kuhn, G. A1 - Diekmann, Bernhard T1 - Late Quaternary vegetation and lake system dynamics in north-eastern Siberia: Implications for seasonal climate variability JF - Quaternary science reviews : the international multidisciplinary research and review journal N2 - Although the climate development over the Holocene in the Northern Hemisphere is well known, palaeolimnological climate reconstructions reveal spatiotemporal variability in northern Eurasia. Here we present a multi-proxy study from north-eastern Siberia combining sediment geochemistry, and diatom and pollen data from lake-sediment cores covering the last 38,000 cal. years. Our results show major changes in pyrite content and fragilarioid diatom species distributions, indicating prolonged seasonal lake-ice cover between similar to 13,500 and similar to 8900 cal. years BP and possibly during the 8200 cal. years BP cold event. A pollen-based climate reconstruction generated a mean July temperature of 17.8 degrees C during the Holocene Thermal Maximum (HTM) between similar to 8900 and similar to 4500 cal. years BP. Naviculoid diatoms appear in the late Holocene indicating a shortening of the seasonal ice cover that continues today. Our results reveal a strong correlation between the applied terrestrial and aquatic indicators and natural seasonal climate dynamics in the Holocene. Planktonic diatoms show a strong response to changes in the lake ecosystem due to recent climate warming in the Anthropocene. We assess other palaeolimnological studies to infer the spatiotemporal pattern of the HTM and affirm that the timing of its onset, a difference of up to 3000 years from north to south, can be well explained by climatic teleconnections. The westerlies brought cold air to this part of Siberia until the Laurentide ice sheet vanished 7000 years ago. The apparent delayed ending of the HTM in the central Siberian record can be ascribed to the exceedance of ecological thresholds trailing behind increases in winter temperatures and decreases in contrast in insolation between seasons during the mid to late Holocene as well as lacking differentiation between summer and winter trends in paleolimnological reconstructions. (C) 2015 Elsevier Ltd. All rights reserved. KW - Diatoms KW - Pollen KW - Summer and winter temperature KW - Holocene Thermal Maximum KW - Aquatic and terrestrial ecosystems KW - Lake-ice cover Y1 - 2016 U6 - https://doi.org/10.1016/j.quascirev.2015.08.014 SN - 0277-3791 VL - 147 SP - 406 EP - 421 PB - Elsevier CY - Oxford ER - TY - JOUR A1 - Brügger, Sandra Olivia A1 - Gobet, Erika A1 - Sigl, Michael A1 - Osmont, Dimitri A1 - Papina, Tatyana A1 - Rudaya, Natalia A1 - Schwikowski-Gigar, Margit A1 - Tinner, Willy T1 - Ice records provide new insights into climatic vulnerability of Central Asian forest and steppe communities JF - Global and planetary change N2 - Forest and steppe communities in the Altai region of Central Asia are threatened by changing climate and anthropogenic pressure. Specifically, increasing drought and grazing pressure may cause collapses of moisture-demanding plant communities, particularly forests. Knowledge about past vegetation and fire responses to climate and land use changes may help anticipating future ecosystem risks, given that it has the potential to disclose mechanisms and processes that govern ecosystem vulnerability. We present a unique paleoecological record from the high-alpine Tsambagarav glacier in the Mongolian Altai that provides novel large-scale information on vegetation, fire and pollution with an exceptional temporal resolution and precision. Our palynological record identifies several late-Holocene boreal forest expansions, contractions and subsequent recoveries. Maximum forest expansions occurred at 3000-2800 BC, 2400-2100 BC, and 1900-1800 BC. After 1800 BC mixed boreal forest communities irrecoverably declined. Fires reached a maximum at 1600 BC, 200 years after the final forest collapse. Our multiproxy data suggest that burning peaked in response to dead biomass accumulation resulting from forest diebacks. Vegetation and fire regimes partly decoupled from climate after 1700 AD, when atmospheric industrial pollution began, and land use intensified. We conclude that moisture availability was more important than temperature for past vegetation dynamics, in particular for forest loss and steppe expansion. The past Mongolian Altai evidence implies that in the future forests of the Russian Altai may collapse in response to reduced moisture availability. KW - Boreal forest diebacks KW - Climatic tipping points KW - Diversity KW - Ice core KW - Moisture change KW - Pollen KW - Microscopic charcoal KW - SCP Y1 - 2018 U6 - https://doi.org/10.1016/j.gloplacha.2018.07.010 SN - 0921-8181 SN - 1872-6364 VL - 169 SP - 188 EP - 201 PB - Elsevier CY - Amsterdam ER - TY - JOUR A1 - Cao, Xianyong A1 - Tian, Fang A1 - Dallmeyer, Anne A1 - Herzschuh, Ulrike T1 - Northern Hemisphere biome changes (> 30 degrees N) since 40 cal ka BP and their driving factors inferred from model-data comparisons JF - Quaternary science reviews : the international multidisciplinary research and review journal N2 - Ongoing and past biome transitions are generally assigned to climate and atmospheric changes (e.g. temperature, precipitation, CO2), but the major regional factors or factor combinations that drive vegetation change often remain unknown. Modelling studies applying ensemble runs can help to partition the effects of the different drivers. Such studies require careful validation with observational data. In this study, fossil pollen records from 741 sites in Europe, 728 sites in North America, and 418 sites in Asia (extracted from terrestrial archives including lake sediments) are used to reconstruct biomes at selected time slices between 40 cal ka BP (calibrated thousand years before present) and today. These results are used to validate Northern Hemisphere biome distributions (>30 degrees N) simulated by the biome model BIOME4 that has been forced with climate data simulated by a General Circulation model. Quantitative comparisons between pollen- and model-based results show a generally good fit at a broad spatial scale. Mismatches occur in central-arid Asia with a broader extent of grassland throughout the last 40 ka (likely due to the over-representation of Artemisia and Chenopodiaceae pollen) and in Europe with over-estimation of tundra at 0 cal ka BP (likely due to human impacts to some extent). Sensitivity analysis reveals that broad-scale biome changes follow the global signal of major postglacial temperature change, although the climatic variables vary in their regional and temporal importance. Temperature is the dominant variable in Europe and other rather maritime areas for biome changes between 21 and 14 ka, while precipitation is highly important in the arid inland regions of Asia and North America. The ecophysiological effect of changes in the atmospheric CO2-concentration has the highest impact during this transition than in other intervals. With respect to modern vegetation in the course of global warming, our findings imply that vegetation change in the Northern Hemisphere may be strongly limited by effective moisture changes, i.e. the combined effect of temperature and precipitation, particularly in inland areas. (C) 2019 Elsevier Ltd. All rights reserved. KW - Biomisation KW - Climate warming KW - Europe KW - Holocene KW - Model-data comparison KW - Northern Asia KW - North America KW - Pollen KW - Siberia KW - Vegetation driver Y1 - 2019 U6 - https://doi.org/10.1016/j.quascirev.2019.07.034 SN - 0277-3791 VL - 220 SP - 291 EP - 309 PB - Elsevier CY - Oxford ER - TY - JOUR A1 - Fritz, Michael A1 - Herzschuh, Ulrike A1 - Wetterich, Sebastian A1 - Lantuit, Hugues A1 - De Pascale, Gregory P. A1 - Pollard, Wayne H. A1 - Schirrmeister, Lutz T1 - Late glacial and holocene sedimentation, vegetation, and climate history from easternmost Beringia (northern Yukon Territory, Canada) JF - Quaternary research : an interdisciplinary journal N2 - Beringian climate and environmental history are poorly characterized at its easternmost edge. Lake sediments from the northern Yukon Territory have recorded sedimentation, vegetation, summer temperature and precipitation changes since similar to 16 cal ka BP. Herb-dominated tundra persisted until similar to 14.7 cal ka BP with mean July air temperatures <= 5 degrees C colder and annual precipitation 50 to 120 mm lower than today. Temperatures rapidly increased during the Bolling/Allerod interstadial towards modern conditions, favoring establishment of Betula-Salix shrub tundra. Pollen-inferred temperature reconstructions recorded a pronounced Younger Dryas stadial in east Beringia with a temperature drop of similar to 1.5 degrees C (similar to 2.5 to 3.0 degrees C below modern conditions) and low net precipitation (90 to 170 mm) but show little evidence of an early Holocene thermal maximum in the pollen record. Sustained low net precipitation and increased evaporation during early Holocene warming suggest a moisture-limited spread of vegetation and an obscured summer temperature maximum. Northern Yukon Holocene moisture availability increased in response to a retreating Laurentide Ice Sheet, postglacial sea level rise, and decreasing summer insolation that in turn led to establishment of Alnus-Berula shrub tundra from similar to 5 cal ka BP until present, and conversion of a continental climate into a coastal-maritime climate near the Beaufort Sea. KW - Trout Lake KW - Laurentide Ice Sheet KW - Younger Dryas KW - Holocene thermal maximum KW - Lake sediments KW - Pollen KW - Temperature reconstruction KW - Precipitation reconstruction KW - WAPLS KW - Modern analogue technique Y1 - 2012 U6 - https://doi.org/10.1016/j.yqres.2012.07.007 SN - 0033-5894 VL - 78 IS - 3 SP - 549 EP - 560 PB - Elsevier CY - San Diego ER - TY - JOUR A1 - Fritz, Michael A1 - Wolter, Juliane A1 - Rudaya, Natalia A1 - Palagushkina, Olga A1 - Nazarova, Larisa B. A1 - Obu, Jaroslav A1 - Rethemeyer, Janet A1 - Lantuit, Hugues A1 - Wetterich, Sebastian T1 - Holocene ice-wedge polygon development in northern Yukon permafrost peatlands (Canada) JF - Quaternary science reviews : the international multidisciplinary research and review journal N2 - Ice-wedge polygon (IWP) peatlands in the Arctic and Subarctic are extremely vulnerable to climatic and environmental change. We present the results of a multidisciplinary paleoenvironmental study on IWPs in the northern Yukon, Canada. High-resolution laboratory analyses were carried out on a permafrost core and the overlying seasonally thawed (active) layer, from an IWP located in a drained lake basin on Herschel Island. In relation to 14 Accelerator Mass Spectrometry (AMS) radiocarbon dates spanning the last 5000 years, we report sedimentary data including grain size distribution and biogeochemical parameters (organic carbon, nitrogen, C/N ratio, delta C-13), stable water isotopes (delta O-18, delta D), as well as fossil pollen, plant macrofossil and diatom assemblages. Three sediment units (SUS) correspond to the main stages of deposition (1) in a thermokarst lake (SW : 4950 to 3950 cal yrs BP), (2) during transition from lacustrine to palustrine conditions after lake drainage (SU2: 3950 to 3120 cal yrs BP), and (3) in palustrine conditions of the IWP field that developed after drainage (SU3: 3120 cal yrs BP to 2012 CE). The lacustrine phase (pre 3950 cal yrs BP) is characterized by planktonic-benthic and pioneer diatom species indicating circumneutral waters, and very few plant macrofossils. The pollen record has captured a regional signal of relatively stable vegetation composition and climate for the lacustrine stage of the record until 3950 cal yrs BP. Palustrine conditions with benthic and acidophilic diatom species characterize the peaty shallow-water environments of the low-centered IWP. The transition from lacustrine to palustrine conditions was accompanied by acidification and rapid revegetation of the lake bottom within about 100 years. Since the palustrine phase we consider the pollen record as a local vegetation proxy dominated by the plant communities growing in the IWP. Ice-wedge cracking in water-saturated sediments started immediately after lake drainage at about 3950 cal yrs BP and led to the formation of an IWP mire. Permafrost aggradation through downward closed-system freezing of the lake talik is indicated by the stable water isotope record. The originally submerged IWP center underwent gradual drying during the past 2000 years. This study highlights the sensitivity of permafrost landscapes to climate and environmental change throughout the Holocene. (C) 2016 Elsevier Ltd. All rights reserved. KW - Permafrost peatlands KW - Arctic KW - Thermokarst KW - Talik KW - Ice-wedge polygon KW - Pollen KW - Diatoms KW - Plant macrofossils KW - Stable water isotopes KW - Deuterium excess Y1 - 2016 U6 - https://doi.org/10.1016/j.quascirev.2016.02.008 SN - 0277-3791 VL - 147 SP - 279 EP - 297 PB - Elsevier CY - Oxford ER - TY - JOUR A1 - Herzschuh, Ulrike A1 - Ni, Jian A1 - Birks, H. John B. A1 - Böhner, Jürgen T1 - Driving forces of mid-Holocene vegetation shifts on the upper Tibetan Plateau, with emphasis on changes in atmospheric CO2 concentrations JF - Quaternary science reviews : the international multidisciplinary research and review journal N2 - Numerous pollen records across the upper Tibetan Plateau indicate that in the early part of the mid-Holocene, Kobresia-rich high-alpine meadows invaded areas formerly dominated by alpine steppe vegetation rich in Artemisia. We examine climate, land-use, and CO2 concentration changes as potential drivers for this marked vegetation change. The climatic implications of these vegetational shifts are explored by applying a newly developed pollen-based moisture-balance transfer-function to fossil pollen spectra from Koucha Lake on the north-eastern Tibetan Plateau (34.0 degrees N; 97.2 degrees E; 4540 m a.s.l.) and Xuguo Lake on the central Tibetan Plateau (31.97 degrees N; 90.3 degrees E; 4595 m a.s.l.), both located in the meadow-steppe transition zone. Reconstructed moisture-balances were markedly reduced (by similar to 150-180 mm) during the early mid-Holocene compared to the late-Holocene. These findings contradict most other records from the Indian monsoonal realm and also most non-pollen records from the Tibetan Plateau that indicate a rather wet early- and mid-Holocene. The extent and timing of anthropogenic land-use involving grazing by large herbivores on the upper Tibetan Plateau and its possible impacts on high-alpine vegetation are still mostly unknown due to the lack of relevant archaeological evidence. Arguments against a mainly anthropogenic origin of Kobresia high-alpine meadows are the discovery of the widespread expansion of obviously 'natural' Kobresia meadows on the south-eastern Tibetan Plateau during the Lateglacial period indicating the natural origin of this vegetation type and the lack of any concurrence between modern human-driven vegetation shifts and the mid-Holocene compositional changes. Vegetation types are known to respond to atmospheric CO2 concentration changes, at least on glacial-interglacial scales. This assumption is confirmed by our sensitivity study where we model Tibetan vegetation at different CO2 concentrations of 375 (present-day), 260 (early Holocene), and 650 ppm (future scenario) using the BIOME4 global vegetation model. Previous experimental studies confirm that vegetation growing on dry and high sites is particularly sensitive to CO2 changes. Here we propose that the replacement of drought-resistant alpine steppes (that are well adapted to low CO2 concentrations) by mesic Kobresia meadows can, at least, be partly interpreted as a response to the increase of CO2 concentration since 7000 years ago due to fertilization and water-saving effects. Our hypothesis is corroborated by former CO2 fertilization experiments performed on various dry grasslands and by the strong recent expansion of high-alpine meadows documented by remote sensing studies in response to recent CO2 increases. KW - Tibetan Plateau KW - Pollen KW - Holocene KW - Transfer function KW - Kobresia meadow KW - Atmospheric CO2 concentration Y1 - 2011 U6 - https://doi.org/10.1016/j.quascirev.2011.03.007 SN - 0277-3791 VL - 30 IS - 15-16 SP - 1907 EP - 1917 PB - Elsevier CY - Oxford ER - TY - JOUR A1 - Hoff, Ulrike A1 - Biskaborn, Boris A1 - Dirksen, Veronika G. A1 - Dirksen, Oleg A1 - Kuhn, Gerhard A1 - Meyer, Hanno A1 - Nazarova, Larisa B. A1 - Roth, Alexandra A1 - Diekmann, Bernhard T1 - Holocene environment of Central Kamchatka, Russia: Implications from a multi-proxy record of Two-Yurts Lake JF - Global and planetary change N2 - Within the scope of Russian German palaeoenvironmental research, Two-Yurts Lake (TYL, Dvuh-Yurtochnoe in Russian) was chosen as the main scientific target area to decipher Holocene climate variability on Kamchatka. The 5 x 2 km large and 26 m deep lake is of proglacial origin and situated on the eastern flank of Sredinny Ridge at the northwestern end of the Central Kamchatka Valley, outside the direct influence of active volcanism. Here, we present results of a multi-proxy study on sediment cores, spanning about the last 7000 years. The general tenor of the TYL record is an increase in continentality and winter snow cover in conjunction with a decrease in temperature, humidity, and biological productivity after 5000-4500 cal yrs BP, inferred from pollen and diatom data and the isotopic composition of organic carbon. The TYL proxy data also show that the late Holocene was punctuated by two colder spells, roughly between 4500 and 3500 cal yrs BP and between 1000 and 200 cal yrs BP, as local expressions of the Neoglacial and Little Ice Age, respectively. These environmental changes can be regarded as direct and indirect responses to climate change, as also demonstrated by other records in the regional terrestrial and marine realm. Long-term climate deterioration was driven by decreasing insolation, while the short-term climate excursions are best explained by local climatic processes. The latter affect the configuration of atmospheric pressure systems that control the sources as well as the temperature and moisture of air masses reaching Kamchatka. (C) 2015 Elsevier B.V. All rights reserved. KW - Kamchatka KW - North Pacific KW - Holocene Climate KW - Palaeolimnology KW - Diatoms KW - Pollen Y1 - 2015 U6 - https://doi.org/10.1016/j.gloplacha.2015.07.011 SN - 0921-8181 SN - 1872-6364 VL - 134 SP - 101 EP - 117 PB - Elsevier CY - Amsterdam ER - TY - JOUR A1 - Li, Huashu A1 - Liu, Xingqi A1 - Herzschuh, Ulrike A1 - Cao, Xianyong A1 - Yu, Zhitong A1 - Wang, Yong T1 - Vegetation and climate changes since the middle MIS 3 inferred from a Wulagai Lake pollen record, Inner Mongolia, Northeastern China JF - Review of palaeobotany and palynology : an international journal N2 - The climate conditions during Marine Isotope Stage (MIS) 3 were similar to present-day conditions, but whether humidity then exceeded present levels is debated, and the driving mechanisms of palaeoclimate evolution since MIS 3 remain unclear. Here, we use pollen data from Wulagai Lake, Inner Mongolia, to reconstruct vegetation and climate changes since the middle MIS 3. The steppe biome is reconstructed as the first dominant biome and the desert biome as the second, and the results show that the vegetation was steppe over the last 43,800 years. Poaceae, Artemisia, Caryophyllaceae and Humulus were abundant from middle to late MIS 3, indicating humid climate conditions. As drought-tolerant species such as Hippophae, Nitraria and Chenopodiaceae spread during MIS 2, the climate became arid. The Holocene is characterized by the dominance of steppe with mixed coniferous-broadleaved forests in the Greater Hinggan Range, and the desert biome retains high affinity scores, indicating that the climate was semi-arid. The climate from middle to late MIS 3 was wetter than in the Holocene; this shift was related to changes in the Northern Hemisphere's solar insolation and ice volume. The humid conditions during MIS 3 were attributed to strong ice–albedo feedback, which led to evaporation that was less than the precipitation. The enhanced evaporation caused by increased solar insolation and decreased ice volume might have exceeded the precipitation during the Holocene and resulted in low effective humidity in the Wulagai Lake basin. KW - Pollen KW - Biome KW - Ice volume KW - Solar insolation Y1 - 2018 U6 - https://doi.org/10.1016/j.revpalbo.2018.12.006 SN - 0034-6667 SN - 1879-0615 VL - 262 SP - 44 EP - 51 PB - Elsevier CY - Amsterdam ER - TY - JOUR A1 - Marquer, Laurent A1 - Gaillard, Marie-Jose A1 - Sugita, Shinya A1 - Poska, Anneli A1 - Trondman, Anna-Kari A1 - Mazier, Florence A1 - Nielsen, Anne Birgitte A1 - Fyfe, Ralph M. A1 - Jonsson, Anna Maria A1 - Smith, Benjamin A1 - Kaplan, Jed O. A1 - Alenius, Teija A1 - Birks, H. John B. A1 - Bjune, Anne E. A1 - Christiansen, Jorg A1 - Dodson, John A1 - Edwards, Kevin J. A1 - Giesecke, Thomas A1 - Herzschuh, Ulrike A1 - Kangur, Mihkel A1 - Koff, Tiiu A1 - Latalowa, Maligorzata A1 - Lechterbeck, Jutta A1 - Olofsson, Jorgen A1 - Seppa, Heikki T1 - Quantifying the effects of land use and climate on Holocene vegetation in Europe JF - Quaternary science reviews : the international multidisciplinary research and review journal N2 - Early agriculture can be detected in palaeovegetation records, but quantification of the relative importance of climate and land use in influencing regional vegetation composition since the onset of agriculture is a topic that is rarely addressed. We present a novel approach that combines pollen-based REVEALS estimates of plant cover with climate, anthropogenic land-cover and dynamic vegetation modelling results. This is used to quantify the relative impacts of land use and climate on Holocene vegetation at a sub-continental scale, i.e. northern and western Europe north of the Alps. We use redundancy analysis and variation partitioning to quantify the percentage of variation in vegetation composition explained by the climate and land-use variables, and Monte Carlo permutation tests to assess the statistical significance of each variable. We further use a similarity index to combine pollen based REVEALS estimates with climate-driven dynamic vegetation modelling results. The overall results indicate that climate is the major driver of vegetation when the Holocene is considered as a whole and at the sub-continental scale, although land use is important regionally. Four critical phases of land-use effects on vegetation are identified. The first phase (from 7000 to 6500 BP) corresponds to the early impacts on vegetation of farming and Neolithic forest clearance and to the dominance of climate as a driver of vegetation change. During the second phase (from 4500 to 4000 BP), land use becomes a major control of vegetation. Climate is still the principal driver, although its influence decreases gradually. The third phase (from 2000 to 1500 BP) is characterised by the continued role of climate on vegetation as a consequence of late-Holocene climate shifts and specific climate events that influence vegetation as well as land use. The last phase (from 500 to 350 BP) shows an acceleration of vegetation changes, in particular during the last century, caused by new farming practices and forestry in response to population growth and industrialization. This is a unique signature of anthropogenic impact within the Holocene but European vegetation remains climatically sensitive and thus may continue to respond to ongoing climate change. (C) 2017 Elsevier Ltd. All rights reserved. KW - Climate KW - Holocene KW - Human impact KW - Land use KW - LPJ-GUESS KW - Europe KW - Pollen KW - REVEALS KW - Vegetation composition Y1 - 2017 U6 - https://doi.org/10.1016/j.quascirev.2017.07.001 SN - 0277-3791 VL - 171 SP - 20 EP - 37 PB - Elsevier CY - Oxford ER -