TY  - JOUR
A1  - Plue, Jan
A1  - De Frenne, Pieter
A1  - Acharya, Kamal
A1  - Brunet, Jörg
A1  - Chabrerie, Olivier
A1  - Decocq, Guillaume
A1  - Diekmann, Martin
A1  - Graae, Bente J.
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Kolb, Annette
A1  - Lemke, Isgard
A1  - Liira, Jaan
A1  - Naaf, Tobias
A1  - Verheyen, Kris
A1  - Wulf, Monika
A1  - Cousins, Sara A. O.
T1  - Where does the community start, and where does it end?
BT  - including the seed bank to reassess forest herb layer responses to the environment
JF  - Journal of vegetation science
N2  - QuestionBelow-ground processes are key determinants of above-ground plant population and community dynamics. Still, our understanding of how environmental drivers shape plant communities is mostly based on above-ground diversity patterns, bypassing below-ground plant diversity stored in seed banks. As seed banks may shape above-ground plant communities, we question whether concurrently analysing the above- and below-ground species assemblages may potentially enhance our understanding of community responses to environmental variation. LocationTemperate deciduous forests along a 2000km latitudinal gradient in NW Europe. MethodsHerb layer, seed bank and local environmental data including soil pH, canopy cover, forest cover continuity and time since last canopy disturbance were collected in 129 temperate deciduous forest plots. We quantified herb layer and seed bank diversity per plot and evaluated how environmental variation structured community diversity in the herb layer, seed bank and the combined herb layer-seed bank community. ResultsSeed banks consistently held more plant species than the herb layer. How local plot diversity was partitioned across the herb layer and seed bank was mediated by environmental variation in drivers serving as proxies of light availability. The herb layer and seed bank contained an ever smaller and ever larger share of local diversity, respectively, as both canopy cover and time since last canopy disturbance decreased. Species richness and -diversity of the combined herb layer-seed bank community responded distinctly differently compared to the separate assemblages in response to environmental variation in, e.g. forest cover continuity and canopy cover. ConclusionsThe seed bank is a below-ground diversity reservoir of the herbaceous forest community, which interacts with the herb layer, although constrained by environmental variation in e.g. light availability. The herb layer and seed bank co-exist as a single community by means of the so-called storage effect, resulting in distinct responses to environmental variation not necessarily recorded in the individual herb layer or seed bank assemblages. Thus, concurrently analysing above- and below-ground diversity will improve our ecological understanding of how understorey plant communities respond to environmental variation.
KW  - Above-ground
KW  - Below-ground
KW  - Canopy
KW  - Disturbance
KW  - Diversity
KW  - Light availability
KW  - NWEurope
KW  - Plant community
KW  - Species co-existence
KW  - Storage effect
Y1  - 2017
U6  - https://doi.org/10.1111/jvs.12493
SN  - 1100-9233
SN  - 1654-1103
VL  - 28
IS  - 2
SP  - 424
EP  - 435
PB  - Wiley
CY  - Hoboken
ER  - 
TY  - JOUR
A1  - Vanneste, Thomas
A1  - Valdes, Alicia
A1  - Verheyen, Kris
A1  - Perring, Michael P.
A1  - Bernhardt-Roemermann, Markus
A1  - Andrieu, Emilie
A1  - Brunet, Jorg
A1  - Cousins, Sara A. O.
A1  - Deconchat, Marc
A1  - De Smedt, Pallieter
A1  - Diekmann, Martin
A1  - Ehrmann, Steffen
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Kolb, Annette
A1  - Lenoir, Jonathan
A1  - Liira, Jaan
A1  - Naaf, Tobias
A1  - Paal, Taavi
A1  - Wulf, Monika
A1  - Decocq, Guillaume
A1  - De Frenne, Pieter
T1  - Functional trait variation of forest understorey plant communities across Europe
JF  - Basic and applied ecology : Journal of the Gesellschaft für Ökologie
N2  - Global environmental changes are expected to alter the functional characteristics of understorey herb-layer communities, potentially affecting forest ecosystem functioning. However, little is known about what drives the variability of functional traits in forest understories. Here, we assessed the role of different environmental drivers in shaping the functional trait distribution of understorey herbs in fragmented forests across three spatial scales. We focused on 708 small, deciduous forest patches located in 16 agricultural landscape windows, spanning a 2500-km macroclimatic gradient across the temperate forest biome in Europe. We estimated the relative effect of patch-scale, landscape-scale and macroclimatic variables on the community mean and variation of plant height, specific leaf area and seed mass. Macroclimatic variables (monthly temperature and precipitation extremes) explained the largest proportion of variation in community trait means (on average 77% of the explained variation). In contrast, patch-scale factors dominated in explaining community trait variation (on average 68% of the explained variation). Notably, patch age, size and internal heterogeneity had a positive effect on the community-level variability. Landscape-scale variables explained only a minor part of the variation in both trait distribution properties. The variation explained by shared combinations of the variable groups was generally negligible. These findings highlight the importance of considering multiple spatial scales in predictions of environmental-change effects on the functionality of forest understories. We propose that forest management sustainability could benefit from conserving larger, historically continuous and internally heterogeneous forest patches to maximise ecosystem service diversity in rural landscapes. (C) 2018 Gesellschaft fur Okologie. Published by Elsevier GmbH. All rights reserved.
KW  - Agricultural landscapes
KW  - Biogeography
KW  - Community ecology
KW  - Forest understorey
KW  - Functional trait diversity
KW  - Fragmentation
KW  - Global environmental change
KW  - Landscape connectivity
KW  - Macroclimatic gradient
KW  - Multi-scale analysis
Y1  - 2018
U6  - https://doi.org/10.1016/j.baae.2018.09.004
SN  - 1439-1791
SN  - 1618-0089
VL  - 34
SP  - 1
EP  - 14
PB  - Elsevier GmbH
CY  - München
ER  - 
TY  - JOUR
A1  - Bernhardt-Römermann, Markus
A1  - Baeten, Lander
A1  - Craven, Dylan
A1  - De Frenne, Pieter
A1  - Hedl, Radim
A1  - Lenoir, Jonathan
A1  - Bert, Didier
A1  - Brunet, Jorg
A1  - Chudomelova, Marketa
A1  - Decocq, Guillaume
A1  - Dierschke, Hartmut
A1  - Dirnboeck, Thomas
A1  - Dörfler, Inken
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Hommel, Patrick
A1  - Jaroszewicz, Bogdan
A1  - Keczynski, Andrzej
A1  - Kelly, Daniel L.
A1  - Kirby, Keith J.
A1  - Kopecky, Martin
A1  - Macek, Martin
A1  - Malis, Frantisek
A1  - Mirtl, Michael
A1  - Mitchell, Fraser J. G.
A1  - Naaf, Tobias
A1  - Newman, Miles
A1  - Peterken, George
A1  - Petrik, Petr
A1  - Schmidt, Wolfgang
A1  - Standovar, Tibor
A1  - Toth, Zoltan
A1  - Van Calster, Hans
A1  - Verstraeten, Gorik
A1  - Vladovic, Jozef
A1  - Vild, Ondrej
A1  - Wulf, Monika
A1  - Verheyen, Kris
T1  - Drivers of temporal changes in temperate forest plant diversity vary across spatial scales
JF  - Global change biology
N2  - Global biodiversity is affected by numerous environmental drivers. Yet, the extent to which global environmental changes contribute to changes in local diversity is poorly understood. We investigated biodiversity changes in a meta-analysis of 39 resurvey studies in European temperate forests (3988 vegetation records in total, 17-75years between the two surveys) by assessing the importance of (i) coarse-resolution (i.e., among sites) vs. fine-resolution (i.e., within sites) environmental differences and (ii) changing environmental conditions between surveys. Our results clarify the mechanisms underlying the direction and magnitude of local-scale biodiversity changes. While not detecting any net local diversity loss, we observed considerable among-site variation, partly explained by temporal changes in light availability (a local driver) and density of large herbivores (a regional driver). Furthermore, strong evidence was found that presurvey levels of nitrogen deposition determined subsequent diversity changes. We conclude that models forecasting future biodiversity changes should consider coarse-resolution environmental changes, account for differences in baseline environmental conditions and for local changes in fine-resolution environmental conditions.
KW  - atmospheric nitrogen deposition
KW  - evenness
KW  - forestREplot
KW  - forest management
KW  - game browsing
KW  - Shannon diversity
KW  - spatiotemporal resurvey data
KW  - species richness
Y1  - 2015
U6  - https://doi.org/10.1111/gcb.12993
SN  - 1354-1013
SN  - 1365-2486
VL  - 21
IS  - 10
SP  - 3726
EP  - 3737
PB  - Wiley-Blackwell
CY  - Hoboken
ER  - 
TY  - JOUR
A1  - Lemke, Isgard H.
A1  - Kolb, Annette
A1  - Graae, Bente J.
A1  - De Frenne, Pieter
A1  - Acharya, Kamal P.
A1  - Blandino, Cristina
A1  - Brunet, Jorg
A1  - Chabrerie, Olivier
A1  - Cousins, Sara A. O.
A1  - Decocq, Guillaume
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Liira, Jaan
A1  - Schmucki, Reto
A1  - Shevtsova, Anna
A1  - Verheyen, Kris
A1  - Diekmann, Martin
T1  - Patterns of phenotypic trait variation in two temperate forest herbs along a broad climatic gradient
JF  - Plant ecology : an international journal
N2  - Phenotypic trait variation plays a major role in the response of plants to global environmental change, particularly in species with low migration capabilities and recruitment success. However, little is known about the variation of functional traits within populations and about differences in this variation on larger spatial scales. In a first approach, we therefore related trait expression to climate and local environmental conditions, studying two temperate forest herbs, Milium effusum and Stachys sylvatica, along a similar to 1800-2500 km latitudinal gradient. Within each of 9-10 regions in six European countries, we collected data from six populations of each species and recorded several variables in each region (temperature, precipitation) and population (light availability, soil parameters). For each plant, we measured height, leaf area, specific leaf area, seed mass and the number of seeds and examined environmental effects on within-population trait variation as well as on trait means. Most importantly, trait variation differed both between and within populations. Species, however, differed in their response. Intrapopulation variation in Milium was consistently positively affected by higher mean temperatures and precipitation as well as by more fertile local soil conditions, suggesting that more productive conditions may select for larger phenotypic variation. In Stachys, particularly light availability positively influenced trait variation, whereas local soil conditions had no consistent effects. Generally, our study emphasises that intra-population variation may differ considerably across larger scales-due to phenotypic plasticity and/or underlying genetic diversity-possibly affecting species response to global environmental change.
KW  - Climate change
KW  - Global environmental change
KW  - Milium effusum
KW  - Phenotypic plasticity
KW  - Intraspecific variation
KW  - Stachys sylvatica
Y1  - 2015
U6  - https://doi.org/10.1007/s11258-015-0534-0
SN  - 1385-0237
SN  - 1573-5052
VL  - 216
IS  - 11
SP  - 1523
EP  - 1536
PB  - Springer
CY  - Dordrecht
ER  - 
TY  - JOUR
A1  - Baeten, Lander
A1  - Warton, David I.
A1  - Van Calster, Hans
A1  - De Frenne, Pieter
A1  - Verstraeten, Gorik
A1  - Bonte, Dries
A1  - Bernhardt-Römermann, Markus
A1  - Cornelis, Johnny
A1  - Decocq, Guillaume
A1  - Eriksson, Ove
A1  - Hedl, Radim
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Hommel, Patrick
A1  - Kirby, Keith J.
A1  - Naaf, Tobias
A1  - Petrik, Petr
A1  - Walther, Gian-Reto
A1  - Wulf, Monica
A1  - Verheyen, Kris
T1  - A model-based approach to studying changes in compositional heterogeneity
JF  - Methods in ecology and evolution : an official journal of the British Ecological Society
Y1  - 2014
SN  - 2041-210X
SN  - 2041-2096
VL  - 5
IS  - 2
SP  - 156
EP  - 164
PB  - Wiley-Blackwell
CY  - Hoboken
ER  - 
TY  - JOUR
A1  - De Frenne, Pieter
A1  - Graae, Bente J.
A1  - Kolb, Annette
A1  - Shevtsova, Anna
A1  - Baeten, Lander
A1  - Brunet, Jörg
A1  - Chabrerie, Olivier
A1  - Cousins, Sara A. O.
A1  - Decocq, Guillaume
A1  - Dhondt, Rob
A1  - Diekmann, Martin
A1  - Gruwez, Robert
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Oster, Mathias
A1  - Saguez, Robert
A1  - Stanton, Sharon
A1  - Tack, Wesley
A1  - Vanhellemont, Margot
A1  - Verheyen, Kris
T1  - An intraspecific application of the leaf-height-seed ecology strategy scheme to forest herbs along a latitudinal gradient
JF  - Ecography : pattern and diversity in ecology ; research papers forum
N2  - We measured LHS traits in 41 Anemone nemorosa and 44 Milium effusum populations along a 1900-2300 km latitudinal gradient from N France to N Sweden. We then applied multilevel models to identify the effects of regional (temperature, latitude) and local (soil fertility and acidity, overstorey canopy cover) environmental factors on LHS traits.
 Both species displayed a significant 4% increase in plant height with every degree northward shift (almost a two-fold plant height difference between the southernmost and northernmost populations). Neither seed mass nor SLA showed a significant latitudinal cline. Temperature had a large effect on the three LHS traits of Anemone. Latitude, canopy cover and soil nutrients were related to the SLA and plant height of Milium. None of the investigated variables appeared to be related to the seed mass of Milium.
 The variation in LHS traits indicates that the ecological strategy determined by the position of each population in this three-factor triangle is not constant along the latitudinal gradient. The significant increase in plant height suggests greater competitive abilities for both species in the northernmost populations. We also found that the studied environmental factors affected the LHS traits of the two species on various scales: spring-flowering Anemone was affected more by temperature, whereas early-summer flowering Milium was affected more by local and other latitude-related factors. Finally, previously reported cross-species correlations between LHS traits and latitude were generally unsupported by our within-species approach.
Y1  - 2011
U6  - https://doi.org/10.1111/j.1600-0587.2010.06399.x
SN  - 0906-7590
VL  - 34
IS  - 1
SP  - 132
EP  - 140
PB  - Wiley-Blackwell
CY  - Malden
ER  - 
TY  - JOUR
A1  - De Frenne, Pieter
A1  - Baeten, Lander
A1  - Graae, Bente J.
A1  - Brunet, Jorg
A1  - Wulf, Monika
A1  - Orczewska, Anna
A1  - Kolb, Annette
A1  - Jansen, Ivy
A1  - Jamoneau, Aurelien
A1  - Jacquemyn, Hans
A1  - Hermy, Martin
A1  - Diekmann, Martin
A1  - De Schrijver, An
A1  - De Sanctis, Michele
A1  - Decocq, Guillaume
A1  - Cousins, Sara A. O.
A1  - Verheyen, Kris
T1  - Interregional variation in the floristic recovery of post-agricultural forests
JF  - The journal of ecology
N2  - 1. Worldwide, the floristic composition of temperate forests bears the imprint of past land use for decades to centuries as forests regrow on agricultural land. Many species, however, display significant interregional variation in their ability to (re)colonize post-agricultural forests. This variation in colonization across regions and the underlying factors remain largely unexplored.
 2. We compiled data on 90 species and 812 species x study combinations from 18 studies across Europe that determined species' distribution patterns in ancient (i.e. continuously forested since the first available land use maps) and post-agricultural forests. The recovery rate (RR) of species in each landscape was quantified as the log-response ratio of the percentage occurrence in post-agricultural over ancient forest and related to the species-specific life-history traits and local (soil characteristics and light availability) and regional factors (landscape properties as habitat availability, time available for colonization, and climate).
 3. For the herb species, we demonstrate a strong (interactive) effect of species' life-history traits and forest habitat availability on the RR of post-agricultural forest. In graminoids, however, none of the investigated variables were significantly related to the RR.
 4. The better colonizing species that mainly belonged to the short-lived herbs group showed the largest interregional variability. Their recovery significantly increased with the amount of forest habitat within the landscape, whereas, surprisingly, the time available for colonization, climate, soil characteristics and light availability had no effect.
 5. Synthesis. By analysing 18 independent studies across Europe, we clearly showed for the first time on a continental scale that the recovery of short-lived forest herbs increased with the forest habitat availability in the landscape. Small perennial forest herbs, however, were generally unsuccessful in colonizing post-agricultural forest even in relatively densely forested landscapes. Hence, our results stress the need to avoid ancient forest clearance to preserve the typical woodland flora.
KW  - ancient forest
KW  - colonization capacity
KW  - forest herbs
KW  - functional traits
KW  - habitat fragmentation
KW  - habitat loss
KW  - life-history traits
KW  - meta-analysis
KW  - plant population and community dynamics
KW  - secondary succession
Y1  - 2011
U6  - https://doi.org/10.1111/j.1365-2745.2010.01768.x
SN  - 0022-0477
VL  - 99
IS  - 2
SP  - 600
EP  - 609
PB  - Wiley-Blackwell
CY  - Malden
ER  - 
TY  - JOUR
A1  - De Frenne, P.
A1  - Kolb, Annette
A1  - Graae, Benete Jessen
A1  - Decocq, Guillaume
A1  - Baltora, S.
A1  - De Schrijver, A.
A1  - Brunet, J.
A1  - Chabrerie, Olivier
A1  - Cousins, Sara A. O.
A1  - Dhondt, Rob
A1  - Diekmann, Martin
A1  - Gruwez, R.
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Liira, J.
A1  - Saguez, R.
A1  - Shevtsova, Anna
A1  - Baskin, Carol C.
A1  - Verheyen, Kris
T1  - A latitudinal gradient in seed nutrients of the forest herb Anemone nemorosa
JF  - Plant biology
N2  - The nutrient concentration in seeds determines many aspects of potential success of the sexual reproductive phase of plants, including the seed predation probability, efficiency of seed dispersal and seedling performance. Despite considerable research interest in latitudinal gradients of foliar nutrients, a similar gradient for seeds remains unexplored. We investigated a potential latitudinal gradient in seed nutrient concentrations within the widespread European understorey forest herb Anemone nemorosa L. We sampled seeds of A. nemorosa in 15 populations along a 1900-km long latitudinal gradient at three to seven seed collection dates post-anthesis and investigated the relative effects of growing degree-hours > 5 degrees C, soil characteristics and latitude on seed nutrient concentrations. Seed nitrogen, nitrogen:phosphorus ratio and calcium concentration decreased towards northern latitudes, while carbon:nitrogen ratios increased. When taking differences in growing degree-hours and measured soil characteristics into account and only considering the most mature seeds, the latitudinal decline remained particularly significant for seed nitrogen concentration. We argue that the decline in seed nitrogen concentration can be attributed to northward decreasing seed provisioning due to lower soil nitrogen availability or greater investment in clonal reproduction. This pattern may have large implications for the reproductive performance of this forest herb as the degree of seed provisioning ultimately co-determines seedling survival and reproductive success.
KW  - Collection date
KW  - latitude
KW  - nutrient stoichiometry
KW  - seed nitrogen
KW  - seed predation
KW  - seed provisioning
KW  - sexual reproduction
KW  - wood anemone
Y1  - 2011
U6  - https://doi.org/10.1111/j.1438-8677.2010.00404.x
SN  - 1435-8603
VL  - 13
IS  - 3
SP  - 493
EP  - 501
PB  - Wiley-Blackwell
CY  - Malden
ER  - 
TY  - JOUR
A1  - De Frenne, Pieter
A1  - Brunet, Jorg
A1  - Shevtsova, Anna
A1  - Kolb, Annette
A1  - Graae, Bente J.
A1  - Chabrerie, Olivier
A1  - Cousins, Sara Ao
A1  - Decocq, Guillaume
A1  - De Schrijver, An
A1  - Diekmann, Martin
A1  - Gruwez, Robert
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Nilsson, Christer
A1  - Stanton, Sharon
A1  - Tack, Wesley
A1  - Willaert, Justin
A1  - Verheyen, Kris
T1  - Temperature effects on forest herbs assessed by warming and transplant experiments along a latitudinal gradient
JF  - Global change biology
N2  - Slow-colonizing forest understorey plants are probably not able to rapidly adjust their distribution range following large-scale climate change. Therefore, the acclimation potential to climate change within their actual occupied habitats will likely be key for their short-and long-term persistence. We combined transplant experiments along a latitudinal gradient with open-top chambers to assess the effects of temperature on phenology, growth and reproductive performance of multiple populations of slow-colonizing understorey plants, using the spring flowering geophytic forb Anemone nemorosa and the early summer flowering grass Milium effusum as study species. In both species, emergence time and start of flowering clearly advanced with increasing temperatures. Vegetative growth (plant height, aboveground biomass) and reproductive success (seed mass, seed germination and germinable seed output) of A. nemorosa benefited from higher temperatures. Climate warming may thus increase future competitive ability and colonization rates of this species. Apart from the effects on phenology, growth and reproductive performance of M. effusum generally decreased when transplanted southwards (e. g., plant size and number of individuals decreased towards the south) and was probably more limited by light availability in the south. Specific leaf area of both species increased when transplanted southwards, but decreased with open-top chamber installation in A. nemorosa. In general, individuals of both species transplanted at the home site performed best, suggesting local adaptation. We conclude that contrasting understorey plants may display divergent plasticity in response to changing temperatures which may alter future understorey community dynamics.
KW  - climate change
KW  - common garden experiment
KW  - forest understorey
KW  - latitude
KW  - local adaptation
KW  - open-top chambers
KW  - phenotypic plasticity
KW  - pot experiment
Y1  - 2011
U6  - https://doi.org/10.1111/j.1365-2486.2011.02449.x
SN  - 1354-1013
VL  - 17
IS  - 10
SP  - 3240
EP  - 3253
PB  - Wiley-Blackwell
CY  - Hoboken
ER  - 
TY  - JOUR
A1  - Verheyen, Kris
A1  - Baeten, Lander
A1  - De Frenne, Pieter
A1  - Bernhardt-Römermann, Markus
A1  - Brunet, Jorg
A1  - Cornelis, Johnny
A1  - Decocq, Guillaume
A1  - Dierschke, Hartmut
A1  - Eriksson, Ove
A1  - Hedl, Radim
A1  - Heinken, Thilo
A1  - Hermy, Martin
A1  - Hommel, Patrick
A1  - Kirby, Keith J.
A1  - Naaf, Tobias
A1  - Peterken, George
A1  - Petrik, Petr
A1  - Pfadenhauer, Joerg
A1  - Van Calster, Hans
A1  - Walther, Gian-Reto
A1  - Wulf, Monika
A1  - Verstraeten, Gorik
T1  - Driving factors behind the eutrophication signal in understorey plant communities of deciduous temperate forests
JF  - The journal of ecology
N2  - 1. Atmospheric nitrogen (N) deposition is expected to change forest understorey plant community composition and diversity, but results of experimental addition studies and observational studies are not yet conclusive. A shortcoming of observational studies, which are generally based on resurveys or sampling along large deposition gradients, is the occurrence of temporal or spatial confounding factors.
 2. We were able to assess the contribution of N deposition versus other ecological drivers on forest understorey plant communities by combining a temporal and spatial approach. Data from 1205 (semi-)permanent vegetation plots taken from 23 rigorously selected understorey resurvey studies along a large deposition gradient across deciduous temperate forest in Europe were compiled and related to various local and regional driving factors, including the rate of atmospheric N deposition, the change in large herbivore densities and the change in canopy cover and composition.
 3. Although no directional change in species richness occurred, there was considerable floristic turnover in the understorey plant community and a shift in species composition towards more shade-tolerant and nutrient-demanding species. However, atmospheric N deposition was not important in explaining the observed eutrophication signal. This signal seemed mainly related to a shift towards a denser canopy cover and a changed canopy species composition with a higher share of species with more easily decomposed litter.
 4. Synthesis. Our multi-site approach clearly demonstrates that one should be cautious when drawing conclusions about the impact of atmospheric N deposition based on the interpretation of plant community shifts in single sites or regions due to other, concurrent, ecological changes. Even though the effects of chronically increased N deposition on the forest plant communities are apparently obscured by the effects of canopy changes, the accumulated N might still have a significant impact. However, more research is needed to assess whether this N time bomb will indeed explode when canopies will open up again.
KW  - atmospheric deposition
KW  - determinants of plant community diversity and structure
KW  - Ellenberg indicator values
KW  - forest herbs
KW  - forest management
KW  - large herbivores
KW  - north-western Europe
KW  - resurveys
KW  - (semi-)permanent plots
Y1  - 2012
U6  - https://doi.org/10.1111/j.1365-2745.2011.01928.x
SN  - 0022-0477
VL  - 100
IS  - 2
SP  - 352
EP  - 365
PB  - Wiley-Blackwell
CY  - Hoboken
ER  -